Two new species of snail-eating snakes of the genus Tropidodipsas (Serpentes, Dipsadidae) from southern Mexico, with notes on related species Author Gruenwald, Christoph I. https://orcid.org/0000-0001-5994-555X Biencom Real Estate, Carretera Chapala-Jocotepec # 57 - 1, C. P. 45920, Ajijic, Jalisco, Mexico. & Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico. cgruenwald@switaki.com Author Toribio-Jimenez, Sarahi Centro de Ciencias de Desarrollo Regional, Universidad Autonoma de Guerrero. Calle Pino s / n. Col. El Roble. C. P. 39640. Acapulco, Guerrero, Mexico. Author Montano-Ruvalcaba, Carlos Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico. Author Franz-Chavez, Hector Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico. Author Penaloza-Montano, Miguel A. Facultad de Ciencias Quimico Biologicas, Universidad Autonoma de Guerrero. Avenida Lazaro Cardenas # 88, Ciudad Universitaria Sur, C. P. 39074, Chilpancingo de los Bravo, Guerrero, Mexico. Author Barrera-Nava, Eduardo Y. Facultad de Ciencias Quimico Biologicas, Universidad Autonoma de Guerrero. Avenida Lazaro Cardenas # 88, Ciudad Universitaria Sur, C. P. 39074, Chilpancingo de los Bravo, Guerrero, Mexico. Author Jones, Jason M. Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico. Author Rodriguez, Christopher M. Los Angeles Zoo and Botanical Gardens, 45333 Zoo Drive, Los Angeles, P. C. 90027, California, USA. Author Hughes, India M. Department of Biology, University of South Alabama, 5871 USA Dr. N., Mobile, P. C. 36688, Alabama, USA. Author Strickland, Jason L. https://orcid.org/0000-0002-1927-7259 Department of Biology, University of South Alabama, 5871 USA Dr. N., Mobile, P. C. 36688, Alabama, USA. Author Reyes-Velasco, Jacobo https://orcid.org/0000-0002-8787-9980 Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico. text Herpetozoa 2021 2021-11-11 34 233 257 http://dx.doi.org/10.3897/herpetozoa.34.e69176 journal article http://dx.doi.org/10.3897/herpetozoa.34.e69176 2682-955X-34-233 FFCB4A62CCBA5446923C8F77B4D3A4DF

Table 3. Meristic variation of Tropidodipsas tricolor sp. nov. and Tropidodipsas papavericola sp. nov.

Species	T. tricolor sp. nov.		T. papavericola sp. nov.
Specimen	INIRENA 2800	INIRENA 2798	INIRENA 2801	INIRENA 2802	INIRENA 2803	INIRENA 2804	INIRENA 2810	INIRENA 2805
Sex	Male	Female	Male	Male	Male	Male	Male	Male
Snout-vent Length	210	310	421	366	363	430	258	336
Tail Length	67	93	137	119	113	127	91	112
Total Length	277	403	558	485	476	557	349	448
Head length	8.30	13.62	18.97	15.25	15.83	18.58	13.03	14.92
Anterior Chinshield	Longer than wide	Longer than wide	Longer than wide	Longer than wide	Longer than wide	Longer than wide	Longer than wide	Longer than wide
Posterior Chinshields	Longer than wide	Longer than wide	Longer than wide	Longer than wide	Longer than wide	Longer than wide	Longer than wide	Longer than wide
Gulars	3	3	4	4	5	4	5	5
Infras in contact with 1st CS	5	5	5	4	4	5	4	4
Ventral Scales	183	183	180	180	188	189	182	179
Subcaudal Scales	79 / 79	79 / 79	76	74 left / 74	73 / 73	69	74	73
Pre-oculars (Side)	1. 1	1. 1*	2. 2	2. 2	2. 2	2. 2	2.2	2. 2
Post-oculars (Side)	2. 2	3. 3	2.2	2. 2	2. 2	1. 1	2. 2	2. 2
Anterior Temporals (Side)	1. 1	1. 1	1. 1	1. 1	1. 1	1. 1	1. 1	1. 1
Posterior Temporals (Side)	2. 2	2. 2	2. 2	2. 2	2. 2	2. 2	2. 2	2. 2
Supralabials (Side)	8. 7	7. 8	7. 7	6. 6	6. 7	5. 6	7. 7	6. 6
Infralabials (Side)	8. 8	8. 9	7. 6	6. 6	6. 6	6. 6	7. 7	6. 6
Post-mentals	0	0	0	0	0	0	0	0
Pale Body Bands (Side)	19. 19	19. 20	-	32. 28	34. 35	29. 27	27. 27	25. 25
Pale Tail Bands (Side)	9. 9	8. 8	-	14. 12	-	11. 11	15. 16	13. 13
Black Body Blotches (Side)	19. 19	19. 21	32. 33	30. 31	35. 35	27. 27	29. 28	26. 26
Black Tail Blotches (Side)	8. 8	8. 8	14. 14	15. 15	-	10. 11	16. 14	13. 13
Total Pale Body Blotch Count (Body & Tail)	28. 28	28. 28	46. 47	45. 46	42. 42	37. 38	45. 42	39. 39
TL/TotL	0.24	0.23	0.25	0.25	0.24	0.23	0.26	0.25
TL/SVL	0.32	0.30	0.33	0.33	0.31	0.30	0.35	0.33
Internasal Length	1.00	2.34	1.70	1.34	1.50	1.46	0.90	1.44
Internasal Width	1.60	2.62	4.48	3.53	3.58	4.74	2.96	3.52
Loreal Length	0.9 / 1.0	1.19 / 1.10	1.73 / 1.70	1.46 / 1.38	1.36 / 1.34	1.26 / 1.22	1.0 / 1.0	1.40 / 1.38
Loreal Width	1.0 / 1.0	1.24 / 1.12	1.84 / 1.82	1.45 / 1.2	1.38 / 1.42	1.40 / 1.46	0.92 / 0.94	1.30 / 1.26
Loreal into orbit	No	No	No	No	No	No	No	No
Pre-frontal Length	1.70	2.34	3.39	2.81	2.89	3.26	2.48	2.36
Pre-frontal Width	2.00	2.64	5.75	4.69	4.69	5.06	3.84	4.33
Prefrontal into orbit	Yes	Yes	No	No	No	No	No	No
Frontal Length	2.80	2.88	4.23	3.58	3.71	3.93	3.31	3.60
Frontal Width	2.40	2.70	3.81	2.73	3.35	3.35	2.71	3.08
Parietal Length	4.30	5.54	6.58	6.82	5.51	6.26	4.61	5.28
Length Anterior Chinshields	2.4 / 2.6	3.49 / 3.63	4.73	4.03	4.31	5.72	3.33	3.58
Length Posterior Chinshields	1.9 / 1.8	2.28 / 2.42	3.72	1.94	2.25	2.67	1.81	2.15
Eye Diameter	2.10	2.80	3.20	3.02	3.11	3.56	2.71	2.85
ED/HL	0.25	0.21	0.17	0.20	0.20	0.19	0.21	0.19
Pre-Frontal Length/Width	0.85	0.89	0.59	0.60	0.62	0.64	0.65	0.55
HL/SVL	0.04	0.04	0.05	0.04	0.04	0.04	0.05	0.04
Midbody scale rows	15	15	15	15	15	15	15	15
Head scale rows	15	15	15	15	15	15	15	15
Caudal scale rows	15	15	15	15	15	15	15	15
Lateral compresion	Yes - Extreme	Yes - Extreme	Yes - Oval	Yes - Oval	Yes - Oval	Yes - Oval	Yes - Oval	Yes - Oval
Anal plate	Undivided	Undivided	Undivided	Undivided	Undivided	Undivided	Undivided	Undivided
Penultimate supralabial enlarged	Yes	Yes	Yes	Yes	Yes	Fused	Yes	Yes
First infraliabals in contact	Yes	Yes	Yes	Yes	Yes	Yes	Yes	Yes
Pale nuchal band	Yes	Yes	Yes	Yes	Yes	Yes	Yes	Yes