Morphological and genetic analysis of Sulawesi Rhacophorus and description of a fourth species endemic to the island Author Hamidy, Amir Laboratory of Herpetology, Museum Zoologicum Bogoriense, Research Centre for Biosystematics and Evolution, National National Research and Innovation Agency (BRIN), Gd. Widyasatwaloka, KST Soekarno, Jl. Raya Jakarta Bogor km 46, Cibinong, Bogor, West Java 16912, Indonesia. Author Riyanto, Awal Laboratory of Herpetology, Museum Zoologicum Bogoriense, Research Centre for Biosystematics and Evolution, National National Research and Innovation Agency (BRIN), Gd. Widyasatwaloka, KST Soekarno, Jl. Raya Jakarta Bogor km 46, Cibinong, Bogor, West Java 16912, Indonesia. Author Munir, Misbahul † Deceased 28 February 2023. Author Gonggoli, Ade Damara Laboratory of Herpetology, Museum Zoologicum Bogoriense, Research Centre for Biosystematics and Evolution, National National Research and Innovation Agency (BRIN), Gd. Widyasatwaloka, KST Soekarno, Jl. Raya Jakarta Bogor km 46, Cibinong, Bogor, West Java 16912, Indonesia. & Graduate Program of Animal Bioscience, Department of Biology, Faculty of Mathematics and Natural Sciences, Bogor Agricultural University, Bogor, West Java 16680, Indonesia. Author Trilaksono, Wahyu Laboratory of Herpetology, Museum Zoologicum Bogoriense, Research Centre for Biosystematics and Evolution, National National Research and Innovation Agency (BRIN), Gd. Widyasatwaloka, KST Soekarno, Jl. Raya Jakarta Bogor km 46, Cibinong, Bogor, West Java 16912, Indonesia. Author Mcguire, Jimmy A. Department of Integrative Biology, University of California, Berkeley, Valley Life Sciences Building, California, 3101, United States of America. & Museum of Vertebrate Zoology, University of California, Berkeley, Valley Life Sciences Building, California, 3101, United States of America. text Zootaxa 2025 2025-01-14 5569 2 201 230 https://doi.org/10.11646/zootaxa.5569.2.1 journal article 307796 10.11646/zootaxa.5569.2.1 73117c8d-abd2-4253-b9fc-358e7a002741 1175-5326 14704152 26E18D07-6CC3-46B2-8DB7-F38296F72C6C

TABLE 2. Morphological variations of adult of Sulawesian Rhacophorus, measurements in (mm). Abbreviations defined in the text.

Characters	Rhacophorus sp.					Rhacophorus monticola				Rhacophorus edentulus				Rhacophorus georgii
	Holotype	Male (n=12)		Female (n=9)		Male (n=10)		Female (n=3)		Male (n=10)		Female (n=11)		Male (n=4)		Female (n=3)
	MZB
	Amph
	31070	Mean ± SD Range		Mean ± SD Range		Mean ± SD Range		Mean ± SD Range		Mean ± SD Range		Mean ± SD Range		Mean ± SD Range		Mean ± SD ange
SVL	42.0	42.6±1.4	40.4–44.6	51.5±2.1	48.1–54.2	35.5±2.7	32.5–40.6	45.5±3.4	43.2–49.5	31.3±1.5	27.8–32.7	35.6±2	31.8–38.1	59.4±5.1	54.6–64.1	71.9±2.6	68.9–73.8
HL	15.4	15.2±0.4	14.5–15.9	16.2±2.7	12.6–18.7	13.2±0.7	12.3–14.7	16±1.3	14.6–17.2	11.6±0.3	11.2–12.1	12.9±0.4	12.4–13.9	23.5±2.3	21.4–26.4	28.9±1.1	27.8–30.0
HW	13.9	13.0±0.8	12.1–14.7	16.7±0.7	15.3–17.6	12.0±0.5	11.2–12.6	15.5±1.5	14.2–17.1	9.9±3.13	11.1–11.7	12.4±0.7	10.8–13.4	23.8±3.2	21–27.3	30.2±1.5	28.7–31.7
SL	3.2	3.1±0.4	2.0–3.6	2.9±0.3	2.3–3.3	2.4±0.4	1.4–2.8	3.5±0.5	3–4	1.7±0.2	1.3–2.0	2.4±0.3	1.6–2.9	4.8±0.6	4.2–5.6	5.8±0.3	5.5–6
IND	3.1	3.6±0.3	3.1–4.1	4.2±0.4	3.7–5.1	3.2±0.3	2.8–3.7	4.7±0.2	4.5–4.9	2.8±0.3	2.3–3.4	3.5±0.3	2.9–4.2	6.0±0.8	5.4–6.9	7.2±0.2	7.0–7.3
IOD	4.9	4.9±0.2	4.6–5.4	5.6±0.2	5.3–6.0	4.5±0.3	4.1–4.9	5.4±0.7	4.8–6.2	4.3±0.3	3.9–4.8	4.4±0.3	3.9–4.8	9.0±1.8	7.2–11.4	11.6±0.8	10.9–12.5
UEW	4.2	4.1±0.3	3.4–4.6	4.5±0.4	3.9–5.4	3.7±0.4	2.7–4.2	4.1±0.5	3.5–4.4	3.3±0.5	2.7–4.4	3.3±0.3	2.9–3.8	5.7±0.3	5.4–6.1	6.5±0.6	6.1–7.2
EL	6.8	6.5±0.5	5.6–7.1	7.4±0.4	6.7–8.2	6.1±0.3	5.6–6.4	5.9±0.4	5.5–6.2	5.4±0.3	4.8–5.7	5.2±0.3	4.8–5.8	8.2±0.5	7.7–8.9	8.6±0.4	8.3–9.1
ED	6.2	5.6±0.4	5.2–6.3	6.5±0.2	6.2–6.8	5.4±0.4	4.7–5.9	4.9±0.2	4.7–5.1	4.8±0.3	4.3–5.2	4.4±0.3	4.0–4.7	6.4±0.6	5.5–6.9	7.4±0.5	6.8–7.8
TD	2.0	2.3±0.3	2.0–2.9	2.9±0.3	2.3–3.3	1.8±0.2	1.4–2.1	2.5±0.2	2.4–2.8	1.8±0.3	1.4–2.3	1.9±0.2	1.6–2.2	4.1±0.4	3.8–4.6	4.9±0.3	4.7–5.2
TEL	1.1	1.1±0.2	0.8–1.4	1.5±0.3	1.1–2.0	1.1±0.2	0.8–1.3	0.8±0.2	0.6–1	1.3±0.3	0.9–1.6	0.7±0.1	0.5–0.9	1.6±0.2	1.4–1.8	2.1±0.2	1.9–2.2
LAL	22.1	22.3±0.8	21.2–23.7	27.8±1.2	25.6–29.5	17.6±0.8	16.6–19.1	29.0±1.8	27.1–30.5	16.4±0.7	15.4–17.4	23.1±1.7	20.1–25.6	36.8±4.8	33–43.3	45.7±3.0	42.3–47.9
1FL	7.2	7.3±0.3	6.9–8.0	9.9±1.0	8.1–11.0	5.9±0.5	5.4–6.9	7.8±0.9	6.9–8.8	5.2±0.3	4.8–5.7	5.9±0.5	5.2–6.9	10.0±1.3	8.7–11.5	13.6±0.4	13.3–14.1
IPTL	2.8	3.3±0.3	2.8–4.0	3.5±0.4	2.6–4.0	2.4±0.2	2.0–2.6	3.1±0.3	2.8–3.4	1.9±0.2	1.7–2.5	2.4±0.1	2.1–2.6	4.5±0.5	4.1–5.3	5.4±0.1	5.3–5.5
3FDW	2.6	2.7±0.3	2.2–3.1	3.7±0.4	3.3–4.3	2.1±0.2	2.0–2.5	2.9±0.4	2.5–3.3	1.9±0.2	1.7–2.2	2.1±0.2	1.8–2.5	3.9±0.4	3.5–4.4	5.5±0.0	5.4–5.5
4FDW	2.3	2.6±0.3	2.2–3.0	3.1±0.5	2.1–3.6	2.0±0.3	1.5–2.9	2.7±0.4	2.4–3.1	1.8±0.2	1.6–2.1	2.1±0.2	1.7–2.3	3.7±0.4	3.4–4.1	5.2±0.2	4.9–5.3
THIGH	21.2	22.1±0.8	21.2–23.4	26.3±1.1	24.3–27.9	18.0±1.1	16.6–19.7	23.8±0.3	23.5–24.1	17.6±0.8	15.9–18.3	19.9±0.6	18.8–20.6	30.1±3.4	27.0–34.0	37.3±2.2	34.8–38.6
TL	21.3	22.2±0.8	21.3–23.5	27.5±1.2	24.6–29.2	17.4±1.1	15.9–19.3	23.7±0.8	22.8–24.4	17.4±0.8	15.9–18.4	20.3±0.8	18.7–21.6	31.8±4.3	28.3–37.3	40.9±2.7	37.9–42.6
FL	18.5	18.8±0.5	18.3–19.9	24.1±1.7	20.4–25.9	15.4±1.3	13.6–17.4	19.2±1.4	18.1–20.9	13.8±0.6	12.6–14.6	14.8±0.8	13.7–16.3	24.1±4.2	20.7–29.4	31.5±0.9	30.5–32.4
IMTL	1.5	1.6±0.2	1.3–2.0	2±0.2	1.7–2.3	1.1±0.2	0.8–1.5	1.7±0.2	1.6–1.8	1.2±0.2	0.9–1.5	1.5±0.2	1.3–1.9	2.7±0.1	2.5–2.9	3.3±0.5	2.8–3.7
1TOEL	6.8	7.3±0.5	6.7–8.0	9.8±1.2	7.3–10.9	5.8±0.5	5.0–6.5	7.6±0.4	7.1–7.9	4.8±0.5	4.0–5.4	5.7±0.4	5.0–6.3	9.6±1.3	8.4–11.2	13.1±1.1	11.9–14.1
4TDW	1.9	2.0±0.2	1.6–2.3	2.6±0.4	1.8–3.1	1.7±0.1	1.6–1.9	2.4±0.2	2.1–2.5	1.5±0.2	1.2–1.7	1.7±0.1	1.4–1.8	3.0±0.4	2.6–3.6	4.1±0.3	3.8–4.4