On the taxonomic status of Asthenodipsas vertebralis (Boulenger, 1900) (Squamata Pareidae) in Borneo with the description of a new species Author Quah, Evan S. H. 0000-0002-5357-1953 Institute of Tropical Biodiversity and Sustainable Development, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, & Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore. kelvinlim @ nus. edu. sg; https: // orcid. org / 0000 - 0002 - 0638 - 9198 kelvinlim@nus.edu.sg Author Lim, Kelvin K. P. 0000-0002-0638-9198 Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore. kelvinlim @ nus. edu. sg; https: // orcid. org / 0000 - 0002 - 0638 - 9198 kelvinlim@nus.edu.sg Author Grismer, L. Lee 0000-0001-8422-3698 Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA. lgrismer @ lasierra. edu; https: // orcid. org / 0000 - 0001 - 8422 - 3698 lgrismer@lasierra.edu text Zootaxa 2021 2021-03-24 4949 1 24 44 journal article 7428 10.11646/zootaxa.4949.1.2 617a0e17-b4c7-4ac5-b371-73293b34d738 1175-5326 4677338 A4B4DDB2-381A-4C48-A3E6-9268D7E17601

TABLE 4. Comparison of selected characters among Bornean species of Asthenodipsas; A. ingeri sp. nov., A. vertebralis, A. borneensis, A. jamilinaisi, A. stuebingi and Bornean populations of A. laevis. Abbreviations are listed in the Materials and Methods.

Characters	Asthenodipsas ingeri sp. nov.	Asthenodipsas vertebralis	Asthenodipsas borneensis	Asthenodipsas jamilinaisi	Asthenodipsas stuebingi	Asthenodipsas laevis Borneo
	(N=1)	(N=13)	(N=7)	(N=2)	(N=4)	(N = 43)
Dorsal scale rows	15/15/15	15/15/13–15	15/15/15	15/15/15	15/15/15	15/15/13
VeS	+	++	+/- or ++	++	+/-	+/-
PoO	2/2	1–2/1–2	2/2	2/2	1–2/1–2	1–3/1–3
T	2+2/2+2	2(0–1)+(1–2)/ 2+2	2(0–1)+(2–3)/ 2(0–2)+(2–3)	1–2 +2/ 2(0–1)+2	2(+1)+2/ 2(+1)+2	2(0–1)+2/ 2(0–2)+2
SupL (orbit contact)	7/7 (3–5/3–5)	7/7 (3+4 or 3–5 [very rare]/3+4)	7–8/7–8 (3+4/3+4)	6/6–7 (3+4/3+4)	6/6 (3+4/3+4)	6–7/6 (3+4 or 3–5/3+4)
InfL (in contact)	6/7 (1st & 3rd)	6–8/6–8 (1st)	4–7/5–7 (2nd or 3rd)	5–6/5–6 (3rd)	4–6/4–6 (2nd or 3rd)	5–7/5–7 (2nd or 3rd)
SaIM	Present	Present	Present	Present	Present	Present
pIM	2	3	2	2	2	2
Sharp vertebral keel	Present	Present	Present	Present	Present	Absent
Sex	♀	♂ (N=5) ♀ (N=8)	♂ (N=2) ♀ (N=5)	♂	♂ (N=2) ♀ (N=2)	♂ (N=20) ♀ (N=23)
Ven	201	190–202 195–202	167–168 166–179	173–175	165 170–175	150–161 153–167
SubC	53	68–78 63–79	45–48 35–42	52–53	40–47 35–36	41–49 34–42
Max SVL (mm)	715	610 503	388 441	378	414 557	325 345
Max TaL (mm)	103	148 114	77 65	78	81 82	72 51