Three new species of torrent treefrogs (Anura, Hylidae) of the Hyloscirtus bogotensis group from the eastern Andean slopes and the biogeographic history of the genus Author Varela-Jaramillo, Andrea 0000-0002-0092-9915 Museo de Zoología, Escuela de Biología, Facultad de Ciencias Exactas, Naturales y Ambientales, Pontificia Universidad Católica del Ecuador, Av. 12 de Octubre y Roca, Aptdo. 17 - 01 - 2184, Quito, Ecuador & 3 Diversity, Santo Domingo Oe 5 - 71 y Cuba, Quito, Ecuador & Institute of Biology, Molecular Evolution and Systematics of Animals, University of Leipzig, Talstrasse 33, 04103 Leipzig, Germany Author Streicher, Jeffrey W. 0000-0002-3738-4162 Herpetology, Natural History Museum, Cromwell Road, London, SW 7 5 BD, United Kingdom Author Venegas, Pablo J. 0000-0002-6501-4492 Museo de Zoología, Escuela de Biología, Facultad de Ciencias Exactas, Naturales y Ambientales, Pontificia Universidad Católica del Ecuador, Av. 12 de Octubre y Roca, Aptdo. 17 - 01 - 2184, Quito, Ecuador & Rainforest Partnership, 4005 Guadalupe St., Austin, TX 78751, USA & Instituto Peruano de Herpetología (IPH), Augusto Salazar Bondy 136, Urb. Higuereta, Surco, Lima, Peru Author Ron, Santiago R. 0000-0001-6300-9350 Museo de Zoología, Escuela de Biología, Facultad de Ciencias Exactas, Naturales y Ambientales, Pontificia Universidad Católica del Ecuador, Av. 12 de Octubre y Roca, Aptdo. 17 - 01 - 2184, Quito, Ecuador text ZooKeys 2025 2025-03-13 1231 233 292 journal article 308615 10.3897/zookeys.1231.124926 1d579864-205c-4b7b-b651-3c94e0eef5a9 DAE3FB4E-155F-4529-856D-12732CB58D1D

Table 4. Acoustic parameters comparing calls of Hyloscirtus albopunctulatus (Lineage A), H. maycu sp. nov. (Lineage B), H. elbakyanae sp. nov. (Lineage D), H. phyllognathus sensu stricto (Lineage E), H. UCS 2 (Lineage F), H. dispersus sp. nov. (Lineage G) and H. torrenticola from type locality. Mean ± SD is given with range in parentheses. Five calls were analyzed per individual.

Parameter	H. albopunctulatus (n = 7)	H. maycu sp. nov. (n = 1)	H. elbakyanae sp. nov. (n = 2)	H. phyllognathus sensu stricto (n = 1)	UCS 2 (n = 1)	H. dispersus sp. nov. (n = 13)	H. torrenticola sensu stricto (n = 2)
Series of calls	2 –> 70	10–13	4–13	3–4	3	1–8	3–4
Rise time of the call (s)	2.28 ± 0.80 (1.43–3.69)	0.832	0.57 ± 0.05 (0.53–0.60)	0.28	0.17	0.61 ± 0.50 (0.053–2.01)	0.16 ± 0.017 (0.148–0.173)
Call duration (s)	0.051 ± 0.005 (0.045–0.058)	0.053	0.06 ± 0.001 (0.055–0.057)	0.04	0.05	0.11 ± 0.015 (0.082–0.130)	0.03 ± 0.001 (0.026–0.028)
Inter-call duration (s)	0.30 ± 0.06 (0.17–0.35)	0.09	0.12 ± 0.007 (0.11–0.12)	0.06	0.05	0.33 ± 0.044 (0.270–0.440)	0.07 ± 0.007 (0.060–0.070)
Dominant frequency (Hz)	2149.84 ± 137.36 (1903.42–2309.0)	2343.8	2321.29 ± 127.86 (2230.88–2411.7)	2177.0	1851.9	2795.41 ± 138.68 (2573.05–3014.60)	2743.79 ± 48.22 (2709.60–2777.80)
Fundamental frequency (Hz)	1214.12 ± 184.71 (1068.72–1571.90)	1171.90	1184.35 ± 30.48 (1162.8–1205.9)	1974.6	1851.9	2700.63 ± 195.66 (2210.80–2924.20)	2743.75 ± 48.16 (2709.70–2777.80)
Bandwidth of the call (Hz)	378.51 ± 24.98 (344.5–421.9)	515.6	581.4 ± 91.36 (516.8–646)	372.5	416.3	359.06 ± 12.71 (343.1–383.85)	459.0 ± 36.06 (433.50–484.50)