Two new species of megasporangiate Sigillariostrobus Schimper (Sigillariostrobaceae) fructifications from the British Coal Measures Author Thomas, B. A. Author Seyfullah, L. J. text Fossil Record 2016 2015-11-16 19 1 1 9 http://dx.doi.org/10.5194/fr-19-1-2016 journal article 305231 10.5194/fr-19-1-2016 92ec696f-9303-4cda-8b3e-1f03fb70c3ec 2193-0074 10965366

Table 1. Morphological comparison of Sigillariostrobus cones with in situ spores referable to Laevigatisporites glabratus.

	S. tieghemi	S. quadrangularis (Les-	S. czarnockii Bochen-	S. leiosporous Abbott,	Sigillariostrobus
	Zeiller, 1884	quereux) White, 1903	ski, 1936	1963	saltwellensis sp.
					nov.
Cone size	160 mm long	Up to 160 mm long, av-	ca. 180 mm long, 17–	Up to 180 mm long,	At least 86 mm
	25–50 mm	erage width 11.6 mm	25 mm broad	20 mm broad	long, 17–20 mm
	broad				broad
Cone axis		1 mm	7–8 mm	4–7 mm	1.2 mm
Sporophyll	Alternating	In whorls	Helical with sporo-	Alternating verticils	Helical
arrangement	whorls with		phylls lying above one	of 4–5 sporophylls;
	8–10 per whorl		another	3–4 mm between
				verticils
Sporophyll	With prominent	Sporophyll laminae	Sporophylls 20–25 mm	Pedicels: 5–7 mm long,	Laminae ca.
shape	heel	9 mm long	long; upper part	1 mm broad at attach-	6.5 mm long,
			sharply pointed with a	ment point, widening to	spreading out-
			hollowed-out deltoid	4 mm	wards from
			shape, margin ciliate.	Laminae: broadly	cone
				triangular, acuminate,
				5–8 mm long, 4–6 mm
				broad, margin ciliate
Sporangia	None observed	4 mm long, 5 mm high,	4–7 mm long, breadth	4–5 mm long, 2 mm	7 mm long,
		4 mm broad	2–4.5 mm;	high, 4–4.5 mm wide;	2.5 mm high
			contain three tetrads of	contains one tetrad of
			megaspores	megaspores
Peduncle	Acicular leaves	None observed	At least 40 mm long	At least 65 mm long	None observed
	or bracts		and 7–8 mm broad,	and 2–5 mm broad,
	attached to the		uppermost 30 mm with	covered with minute
	upper portion		long, triangular and	stiff spines up to 1 mm
			pointed sterile leaves	long
			25 mm long and 25 mm
			wide at the base
Age and	Age unknown,	Brazil Formation,	Duckmantian, Upper	Upper Freeport (no.	Upper Langsettian–
locality	Valenciennes,	Upper Pottsville,	Silesian Basin, Poland	7) Coal Allegheny,	Duckmantian
	France	Duckmantian/		mid-Pennsylvanian,
		Bolsovian		Bolsovian/Asturian,
		Indiana, USA		southeastern Ohio,
				USA