A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species Author Dewa, Yuna 0000-0002-8512-6782 Graduate School of Agriculture, Forestry and Fisheries, Kagoshima University, 1 - 21 - 24 Korimoto, Kagoshima 890 - 0065, Japan. k2533560@kadai.jp Author Tashiro, Satokuni Yaeyama Field Station, Fisheries Technology Institute, Japan Fisheries Research and Education Agency, 148 Fukai-Ohta, Ishigaki, Okinawa 907 - 0451, Japan. Author Motomura, Hiroyuki 0000-0002-7448-2482 The Kagoshima University Museum, 1 - 21 - 30 Korimoto, Kagoshima 890 - 0065, Japan. text Zootaxa 2023 2023-11-17 5374 3 333 360 https://www.mapress.com/zt/article/download/zootaxa.5374.3.2/52293 journal article 278751 10.11646/zootaxa.5374.3.2 ff275bc8-83fd-4930-8393-4292023a9092 1175-5326 10150204 D8FF43DE-4A2F-4E72-A621-B0A11EEACE72

TABLE 1. Counts and morphometric measurements (as % of SL) of specimens of Enneapterygius olivaceus n. sp.

	Holotype	Paratypes
Means in parentheses
	KAUM–I. 101347	n = 16	n = 11
	Japan	Japan and Philippines
	Male	Males	Females
Standard length (SL; mm)	19.3	14.4–25.2	13.5–22.7
Counts
Dorsal-fin rays	III, XI, 10	III, XI–XIII, 8–10	III, XI–XII, 9–10
Anal-fin rays	I, 16	I, 16–17	I, 16–17
Pectoral-fin rays	iii + 5 + vii = 15	i–v + 3–7 + vii–viii = 14–17	i–v + 3–7 + vi–viii = 14–16
Scale rows in longitudinal series	31	30–32	30–32
Pored lateral-line scales	13	12–14	12–13
Notched lateral-line scales	20	18–22	18–22
Scales above of 1st PLL	2	1–2½	2–3
Scales below 2nd dorsal fin	2½	2–3	2–3
Scales below of 1st NLL	3	2–3	2–3
Circumpeduncular scales	8	8	8
Mandibular pore formula	3 + 1 + 3	3 + 1 + 3	3 + 1 + 3 (rarely 4 + 1 + 4)
Measurements (% SL)
Body depth	20.3	19.3–22.8 (20.6)	17.3–22.1 (19.9)
Body width	20.6	17.4–22.8 (19.3)	16.7–20.4 (18.5)
Head length	28.4	29.2–33.0 (31.4)	28.7–32.3 (31.0)
Snout length	9.4	6.7–11.7 (9.9)	8.6–12.3 (10.0)
Orbit diameter	7.6	8.0–12.1 (9.9)	8.1–11.7 (10.2)
Interorbital width	2.4	1.2–3.4 (2.6)	1.2–2.8 (2.2)
Upper-jaw length	11.4	10.2–13.1 (11.7)	9.5–12.0 (10.9)
Postorbital length	13.7	12.1–15.5 (13.5)	11.6–13.8 (12.8)
Pre-1st-dorsal-fin length	25.2	23.5–28.4 (25.8)	23.9–28.4 (26.3)
Pre-2nd-dorsal-fin length	35.5	33.6–39.3 (36.5)	36.2–39.1 (37.7)
Pre-3rd-dorsal-fin length	69.1	66.8–71.8 (70.0)	69.3–73.3 (70.9)
Pre-anal-fin length	47.8	47.3–52.9 (49.3)	48.3–52.6 (50.9)
Anal-fin base length	39.1	38.8–43.0 (40.7)	36.2–41.3 (39.2)
Pre-pectoral-fin length	32.5	30.3–35.7 (32.4)	31.2–35.2 (33.0)
Pre-pelvic-fin length	24.5	21.9–27.7 (24.1)	21.7–26.6 (23.9)
Caudal peduncle length	13.2	11.0–15.3 (13.3)	10.8–15.1 (12.9)
Caudal peduncle depth	9.9	7.2–9.9 (8.8)	7.7–9.2 (8.5)
1st spine length of 1st dorsal fin	13.8	11.8–15.5 (14.0)	11.2–14.9 (12.7)
2nd spine length of 1st dorsal fin	12.2	10.5–14.7 (12.6)	9.5–13.5 (11.1)
3rd spine length of 1st dorsal fin	9.8	7.3–11.1 (9.4)	6.8–9.3 (8.2)
1st dorsal-fin base length	5.6	4.7–6.8 (5.8)	4.0–6.4 (5.3)
1st spine length of 2nd dorsal fin	12.6	10.0–14.2 (12.9)	11.4–14.0 (12.6)
2nd spine length of 2nd dorsal fin	13.4	12.6–16.0 (14.5)	12.5–14.7 (13.7)
3rd spine length of 2nd dorsal fin	13.7	13.1–16.1 (14.9)	12.8–15.0 (14.1)
2nd dorsal-fin base length	27.5	25.3–33.4 (29.2)	23.9–31.1 (28.3)
1st ray length of 3rd dorsal fin	14.6	11.9–18.5 (16.2)	13.4–18.1 (15.8)
2nd ray length of 3rd dorsal fin	13.7	13.8–18.4 (16.4)	14.2–17.2 (15.6)
3rd ray length of 3rd dorsal fin	13.2	13.2–17.8 (15.0)	12.7–16.1 (15.0)
3rd dorsal fin base length	20.1	15.3–19.0 (17.4)	16.4–19.1 (17.5)
Pectoral-fin length	32.3	30.1–39.4 (34.9)	29.6–39.9 (34.8)
1st ray length of pelvic fin	13.9	14.4–19.1 (16.7)	12.2–18.0 (15.8)
2nd ray length of pelvic fin	22.2	21.8–27.9 (25.1)	20.5–27.3 (24.7)