Phylogeographic pattern and taxonomic revision of the Kaloula baleata species complex (Amphibia, Anura, Microhylidae) with description of two new species from Indochina Author Gorin, Vladislav A. 0000-0002-9017-1502 Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, Moscow 119234, Russia Author Orlov, Nikolai L. 0000-0003-4401-348X Division of Herpetology & Ormithology, Zoological Institute, Russian Academy of Sciences, St. Petersburg 199034 Universitetskaya nab., 1, 33701, Russia Author Bragin, Andrey M. 0000-0002-3621-9763 Joint Vietnam - Russia Tropical Science & Technology Research Center, 63 Nguyen Van Huyen Road, Nghia Do, Cau Giay, Hanoi 122000, Vietnam Author Pawangkhanant, Parinya 0000-0002-0947-5729 Division of Fishery, School of Agriculture & Natural Resources, University of Phayao, Phayao 56000, Thailand & Rabbit in the Moon Foundation, Suanphueng, Ratchaburi 70180, Thailand Author Milto, Konstantin D. 0000-0003-4061-0164 Division of Herpetology & Ormithology, Zoological Institute, Russian Academy of Sciences, St. Petersburg 199034 Universitetskaya nab., 1, 33701, Russia Author Le, Dac Xuan 0000-0003-4586-6214 Joint Vietnam - Russia Tropical Science & Technology Research Center, 63 Nguyen Van Huyen Road, Nghia Do, Cau Giay, Hanoi 122000, Vietnam Author Nguyen, Tan Van 0000-0001-5413-968X Institute for Research & Training in Medicine, Biology & Pharmacy, Duy Tan University, Da Nang 550000, Vietnam Author Dufresnes, Christophe 0000-0002-8497-8908 College of Medicine & Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang 550000, Vietnam Author Suwannapoom, Chatmongkon 0000-0002-3342-1464 Division of Fishery, School of Agriculture & Natural Resources, University of Phayao, Phayao 56000, Thailand Author Poyarkov, Nikolay A. 0000-0002-7576-2283 Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, Moscow 119234, Russia & Joint Vietnam - Russia Tropical Science & Technology Research Center, 63 Nguyen Van Huyen Road, Nghia Do, Cau Giay, Hanoi 122000, Vietnam text Herpetozoa 2024 2024-12-11 37 391 420 journal article 306388 10.3897/herpetozoa.37.e137394 5be396ed-af3c-4af5-b187-1c48444979fb E97A128A-13B5-4781-8FEC-ADC5CB37AB45

Table 2. Uncorrected average interspecific (below and above diagonal) and intraspecific (on the diagonal) genetic p - distances for the 16 S rRNA mtDNA gene fragment (in percentage) are given for species of the genus Kaloula.

	Species	1	2	3	4	5	6	7	8	9	10	11	12	13	14	15	16	17	18	19	20	21	22	23	24
1	Kaloula discordia sp. nov.	0.39	4.45	6.17	5.41	5.40	5.47	6.29	4.33	7.00	6.74	9.57	11.17	8.14	6.25	6.98	7.17	6.08	7.45	7.00	6.65	12.77	9.41	10.61	6.54
2	Kaloula laosensis sp. nov.	4.45	0.22	4.77	6.81	6.59	7.03	7.09	6.28	7.24	8.14	10.74	12.08	7.93	7.65	8.95	8.47	7.47	6.78	7.01	7.89	14.01	10.69	10.64	8.39
3	K. indochinensis	6.17	4.77	0.37	7.24	7.25	6.85	8.20	6.26	7.90	7.45	11.87	12.98	8.37	7.92	8.82	8.70	6.86	6.96	7.29	7.84	15.09	11.44	11.07	7.81
4	K. baleata 1	5.41	6.81	7.24	1.37	2.72	2.08	3.22	4.79	6.64	7.07	9.72	10.85	8.05	6.90	6.71	7.37	6.24	8.19	7.21	6.72	12.46	9.26	11.25	7.57
5	K. baleata 2	5.40	6.59	7.25	2.72	2.56	2.65	3.96	5.36	6.74	7.29	10.26	11.31	7.92	7.35	7.14	7.74	6.76	8.19	7.04	6.82	12.59	9.67	10.59	7.84
6	K. baleata 3	5.47	7.03	6.85	2.08	2.65	0.00	3.00	4.98	6.65	6.44	9.10	10.74	7.57	6.86	6.80	7.06	5.99	7.80	6.51	6.26	13.66	9.00	9.95	7.34
7	K. latidisca	6.29	7.09	8.20	3.22	3.96	3.00	1.45	6.04	7.28	7.47	9.71	11.01	8.89	7.41	7.26	7.74	7.13	8.43	7.82	7.24	14.05	9.41	11.83	7.66
8	K. mediolineata	4.33	6.28	6.26	4.79	5.36	4.98	6.04	0.55	5.89	5.43	8.74	10.49	6.12	5.70	6.95	6.05	5.61	6.81	5.46	5.39	12.30	8.49	10.02	4.97
9	K. borealis	7.00	7.24	7.90	6.64	6.74	6.65	7.28	5.89	0.00	7.43	10.12	11.18	2.07	7.07	7.92	7.46	6.44	2.76	1.93	6.78	13.40	9.91	10.11	7.75
10	K. conjuncta	6.74	8.14	7.45	7.07	7.29	6.44	7.47	5.43	7.43	1.39	8.37	9.78	8.18	5.08	8.59	4.56	3.53	7.70	8.00	3.56	11.65	8.05	7.06	2.57
11	K. kalingensis	9.57	10.74	11.87	9.72	10.26	9.10	9.71	8.74	10.12	8.37	2.07	6.18	10.05	9.39	11.22	8.23	7.55	9.74	9.43	7.52	10.16	4.43	11.20	7.78
12	K. kokacii	11.17	12.08	12.98	10.85	11.31	10.74	11.01	10.49	11.18	9.78	6.18	3.39	11.41	10.64	11.73	9.29	9.26	10.64	10.51	8.25	9.62	4.48	12.17	9.25
13	K. nonggangensis	8.14	7.93	8.37	8.05	7.92	7.57	8.89	6.12	2.07	8.18	10.05	11.41	0.00	7.73	8.60	8.77	8.05	2.76	1.43	8.35	14.14	10.02	11.05	8.14
14	K. picta	6.25	7.65	7.92	6.90	7.35	6.86	7.41	5.70	7.07	5.08	9.39	10.64	7.73	0.31	9.40	3.83	3.15	7.54	8.18	3.51	11.78	8.75	9.18	4.56
15	K. pulchra	6.98	8.95	8.82	6.71	7.14	6.80	7.26	6.95	7.92	8.59	11.22	11.73	8.60	9.40	0.68	8.88	8.98	8.56	7.93	9.27	14.15	10.63	12.93	8.41
16	K. rigida	7.17	8.47	8.70	7.37	7.74	7.06	7.74	6.05	7.46	4.56	8.23	9.29	8.77	3.83	8.88	1.90	2.73	8.56	7.99	2.45	9.90	7.82	7.74	3.58
17	K. cf. rigida	6.08	7.47	6.86	6.24	6.76	5.99	7.13	5.61	6.44	3.53	7.55	9.26	8.05	3.15	8.98	2.73	n / c	7.59	7.45	1.92	10.72	7.78	6.82	3.88
18	K. rugifera	7.45	6.78	6.96	8.19	8.19	7.80	8.43	6.81	2.76	7.70	9.74	10.64	2.76	7.54	8.56	8.56	7.59	0.00	1.98	8.12	14.62	9.33	10.70	7.98
19	K. verrucosa	7.00	7.01	7.29	7.21	7.04	6.51	7.82	5.46	1.93	8.00	9.43	10.51	1.43	8.18	7.93	7.99	7.45	1.98	0.27	7.48	13.70	9.19	10.44	7.68
20	K. walteri	6.65	7.89	7.84	6.72	6.82	6.26	7.24	5.39	6.78	3.56	7.52	8.25	8.35	3.51	9.27	2.45	1.92	8.12	7.48	0.30	9.38	6.72	6.61	3.19
21	Kaloula sp. Panay	12.77	14.01	15.09	12.46	12.59	13.66	14.05	12.30	13.40	11.65	10.16	9.62	14.14	11.78	14.15	9.90	10.72	14.62	13.70	9.38	0.00	8.51	12.80	10.47
22	Kaloula sp. East Luzon	9.41	10.69	11.44	9.26	9.67	9.00	9.41	8.49	9.91	8.05	4.43	4.48	10.02	8.75	10.63	7.82	7.78	9.33	9.19	6.72	8.51	1.85	10.44	7.51
23	Kaloula sp. Samar & Leyte	10.61	10.64	11.07	11.25	10.59	9.95	11.83	10.02	10.11	7.06	11.20	12.17	11.05	9.18	12.93	7.74	6.82	10.70	10.44	6.61	12.80	10.44	1.56	7.78
24	Kaloula sp. Sibuyan	6.54	8.39	7.81	7.57	7.84	7.34	7.66	4.97	7.75	2.57	7.78	9.25	8.14	4.56	8.41	3.58	3.88	7.98	7.68	3.19	10.47	7.51	7.78	1.66