Systematics, distribution and morphology of the newt parasitic water mites of the subgenus Lurchibates Goldschmidt & Fu, 2011 (Acari, Hydrachnidia Hygrobatidae, Hygrobates Koch, 1837), including the description of four new species and a key to all so far known species Author Goldschmidt, Tom Bavarian State Collection of Zoology, Section Arthropoda varia, Münchhausenstrasse 21, D- 81247 Munich, Germany. Author Nishikawa, Kanto 0000-0002-6274-4959 Graduate School of Global Environmental Studies, Kyoto University, Yoshida Hon-machi, Sakyo-ku, Kyoto 606 - 8501, Japan. nishikawa. kanto. 8 v @ kyoto-u. ac. jp; https: // orcid. org / 0000 - 0002 - 6274 - 4959 & Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu-cho, Sakyo-ku, Kyoto 606 - 8501, Japan. Center for Molecular Biodiversity Research, National Museum of Nature and Science, Tsukuba, 305 - 0005 Japan. s-hiruta @ kahaku. go. jp; https: // orcid. org / 0000 - 0001 - 7890 - 1720 nishikawa.kanto.8v@kyoto-u.ac.jp Author Hiruta, Shimpei F. 0000-0001-7890-1720 s-hiruta@kahaku.go.jp Author Pfingstl, Tobias Institute of Biology, University of Graz, Universitätsplatz 2, 8010 Graz, Austria. Author Jiang, Jian-Ping Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China. Author Shimano, Satoshi Science Research Center, Hosei University. Fujimi 2 - 17 - 1, Chiyoda-ku, Tokyo 102 - 8160, Japan. text Zootaxa 2021 2021-06-11 4985 1 1 36 journal article 4938 10.11646/zootaxa.4985.1.1 e8a47bc6-e0ca-4389-9c90-b23a1e56581f 1175-5326 4929035 4EAFC76B-D4E4-4D96-8D0D-1B24E44975C7

Table 1. Overview on the available information on the currently known species of Hygrobates (Lurchibates).

Hygrobates (Lurchibates) species	Known stages / sex	Distribution (based on collecting sites, host distribution)	Newt host species
salamandrarum Goldschmidt, Gerecke & Alberti, 2002	female	eastern China	Pachytriton granulosus Chang, 1933
ancistrophorus Goldschmidt & Koehler, 2007	male, female	northern Laos	Laotriton laoensis (Stuart & Papenfuss, 2002)
forcipifer Goldschmidt & Koehler, 2007	male, female, deutonymph, larva	northern Vietnam	Paramesotriton deloustali (Bourret, 1934)
			P. guangxiensis (Huang, Tang, and Tang, 1983)
aloisii Goldschmidt & Fu, 2011	male, female, deutonymph, larva	Hong Kong	Paramesotriton hongkongensis (Myers and Leviton, 1962)
intermedius Goldschmidt, Nishikawa & Shimano 2020	male, female	southern central China (Jiangxi)	Paramesotriton qixilingensis Yuan, Zhao, Jiang, Hou, He, Murphy & Che, 2014
pilosus Goldschmidt, Nishikawa & Shimano 2020	male, female	southern China (Guangdong)	Paramesotriton yunwuensis Wu, Jiang & Hanken, 2010
robustipalpis Goldschmidt, Nishikawa & Shimano 2020	male, female	southern & southern central China (Guangxi, Guizhou, Guangdong, Hunan)	Pachytriton inexpectatus Nishikawa, Jiang, Matsui & Mo, 2011
			Paramesotriton caudopunctatus (Liu & Hu, 1973)
incognitus sp. nov.	female	southern China (southern Guangxi); northeastern Vietnam (Cao Bang)	Paramesotriton guangxiensis (Huang, Tang, and Tang, 1983)
macrochela sp. nov.	male, female	southern & southern central China (Guangxi, Guizhou, Guangdong, Hunan)	Pachytriton inexpectatus Nishikawa, Jiang, Matsui & Mo, 2011
malosimilis sp. nov.	male	southern & southern central China (Guangxi, Guizhou, Guangdong, Hunan)	Pachytriton inexpectatus Nishikawa, Jiang, Matsui & Mo, 2011
fragmentarius sp. nov.	(only fragments)	southern China (Guangdong)	Paramesotriton yunwuensis Wu, Jiang & Hanken, 2010