Four new species of the Hylomyscus anselli group (Mammalia: Rodentia: Muridae) from the Democratic Republic of Congo and Tanzania Author Kerbis Peterhans, Julian C. 3B4A1A6B-B01E-4FF8-ADF1-6CF1FE102D20 College of Arts & Sciences, Roosevelt University, 430 S Michigan Ave, Chicago, IL USA 60605 & Science and Education, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA & P. O. Box 691, Iringa, Tanzania Author Hutterer, Rainer 16023337-0832-4490-89A9-846AC3925DD8 Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D- 53113 Bonn, Germany Author Doty, Jeffrey B. 33AFA407-C88F-4A8C-B694-AF29E3F894D7 U. S. Centers for Disease Control and Prevention, Poxvirus and Rabies Branch, 1600 Clifton Rd. Atlanta, GA 30333, USA Author Malekani, Jean M. B1445794-1C0A-4861-8AED-1A8C3434BBD3 Département de Biologie, Faculté des Sciences, Université de Kinshasa, Kinshasa, Democratic Republic of Congo Author Moyer, David C. D73C593D-82B0-4587-8C70-52E3AA72B538 Science and Education, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA & P. O. Box 691, Iringa, Tanzania Author Krásová, Jarmila 1F1FFB1B-E594-404B-AE93-EB8D9B9BA101 Department of Zoology, Faculty of Science, University of South Bohemia, České Budějovice, Czech Republic Author Bryja, Josef 63C1A788-1102-4CBC-A0D3-B2434097359D Institute of Vertebrate Biology, Academy of Sciences of the Czech Republic, Brno, Czech Republic & Department of Botany and Zoology, Faculty of Science, Masaryk University, Brno, Czech Republic * Corresponding author: Email: jkerbis @ fieldmuseum. org jkerbis@fieldmuseum.org Author Banasiak, Rebecca A. 5DD1017E-0692-4049-8420-C4705EB2B505 Science and Education, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA Author Demos, Terrence C. 90A9F9F1-8113-4B5E-B0E2-DA0212D118E4 Science and Education, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA text Bonn zoological Bulletin 2020 2020-02-28 69 1 55 83 http://dx.doi.org/10.20363/bzb-2020.69.1.055 journal article 310702 10.20363/BZB-2020.69.1.055 9105b4c6-4c2a-499a-a93e-5524ff2c6c79 2190–7307 15759615 1FD4D09C-D160-4159-A50D-20B6FBC7D9E9

Table 4. Craniodental measurements in millimeters (mean ± 1 SD and range) for eight species in the Hylomyscus anselli group. Abbreviated variables are defined in the text.

Variable	kerbispeterhansi	stanleyi	mpungamachagorum	arcimontensis	thornesmithae	anselli	heinrichorum	pygmaeus
	(n = 51)	(n = 22)	(n = 4)	(n = 53)	(n = 5)	(n = 4)	(n = 13)	(n = 1)
ONL	26.36 ± 0.75	27.16 ± 0.56	25.25 ± 0.08	25.69 ± 0.69	22.84 ± 0.61	25.98 ± 0.30	26.73 ± 0.79	17.45
	(24.68–27.96)	(26.20–28.48)	(25.17–25.36)	(24.12–27.25)	(22.09–23.39)	(25.74–26.39)	(25.52–28.12)
CI	24.73 ± 0.87	24.84 ± 0.56	23.34 ± 0.31	23.80 ± 0.70	21.42 ± 0.47	23.98 ± 0.57	24.77 ± 0.75	15.89
	(22.66–26.53)	(23.60–26.42)	(22.95–23.70)	(22.43–25.61)	(21.06–22.08)	(23.28–24.68)	(23.68–26.18)
ZB	13.19 ± 0.42	13.35 ± 0.25	12.65 ± 0.12	12.85 ± 0.45	11.66 ± 0.10	12.84 ± 0.22	12.99 ± 0.38	9.68
	(12.25–14.19)	(12.86–13.95)	(12.48–12.73)	(11.91–13.89)	(11.55–11.80)	(12.53–13.05)	(12.52–13.70)
BBC	11.81 ± 0.27	12.14 ± 0.24	11.65 ± 0.16	11.66 ± 0.34	11.07 ± 0.49	11.80 ± 0.39	11.81 ± 0.22	9.27
	(11.25–12.51)	(11.62–12.48)	(11.51–11.83)	(10.90–12.47)	(10.64–11.62)	(11.24–12.12)	(11.55–12.29)
BOC	6.10 ± 0.18	6.11 ± 0.12	5.92 ± 0.11	6.12 ± 0.22	5.62 ± 0.17	6.18 ± 0.18	6.26 ± 0.12	4.94
	(5.58–6.45)	(5.93–6.32)	(5.78–6.05)	(5.55–6.54)	(5.42–5.81)	(6.00–6.40)	(6.06–6.43)
IO	4.35 ± 0.10	4.60 ± 0.12	4.52 ± 0.21	4.36 ± 0.15	3.89 ± 0.08	4.41 ± 0.16	4.53 ± 0.08	3.74
	(4.17–4.57)	(4.46–4.91)	(4.34–4.80)	(4.09–4.66)	(3.77–3.99)	(4.20–4.57)	(4.42–4.69)
LN	9.45 ± 0.46	10.14 ± 0.33	8.71 ± 0.07	9.19 ± 0.35	7.69 ± 0.31	9.13 ± 0.27	9.16 ± 0.49	4.66
	(8.44–10.69)	(9.26–10.61)	(8.64–8.81)	(8.44–9.76)	(7.32–7.97)	(8.81–9.38)	(8.06–9.94)
BR	4.56 ± 0.27	4.55 ± 0.14	4.43 ± 0.11	4.45 ± 0.20	4.11 ± 0.20	4.59 ± 0.07	4.35 ± 0.24	3.23
	(4.16–5.60)	(4.27–4.97)	(4.27–4.50)	(4.08–4.96)	(3.94–4.42)	(4.48–4.63)	(4.01–4.73)
PPL	9.03 ± 0.44	9.36 ± 0.27	8.98 ± 0.24	8.96 ± 0.31	8.62 ± 0.33	9.11 ± 0.24	9.40 ± 0.44	6.28
	(8.12–10.32)	(8.75–10.05)	(8.69–9.28)	(8.37–9.63)	(8.23–8.94)	(8.87–9.45)	(8.58–10.20)
LBP	4.50 ± 0.18	4.76 ± 0.18	4.26 ± 0.06	4.42 ± 0.21	4.18 ± 0.14	4.79 ± 0.17	4.45 ± 0.20	2.91
	(3.92–4.81)	(4.36–5.09)	(4.19–4.34)	(3.95–4.83)	(4.00–4.37)	(4.60–4.93)	(4.08–4.77)
LIF	5.59 ± 0.25	5.48 ± 0.16	5.09 ± 0.19	5.38 ± 0.30	4.38 ± 0.22	5.23 ± 0.22	5.84 ± 0.27	2.83
	(5.02–6.06)	(5.29–5.78)	(4.89–5.34)	(4.78–6.11)	(4.10–4.67)	(5.09–5.55)	(5.33–6.33)
LD	7.82 ± 0.32	7.59 ± 0.17	6.95 ± 0.19	7.22 ± 0.25	6.55 ± 0.29	7.19 ± 0.19	7.47 ± 0.35	4.39
	(7.25–8.62)	(7.38–8.03)	(6.74–7.20)	(6.76–7.83)	(6.14–6.82)	(7.03–7.41)	(7.05–8.15)
BZP	2.39 ± 0.12	2.28 ± 0.13	2.30 ± 0.05	2.31 ± 0.12	1.90 ± 0.09	2.34 ± 0.14	2.64 ± 0.17	1.76
	(2.16–2.75)	(2.07–2.57)	(2.24–2.36)	(2.01–2.54)	(1.79–2.00)	(2.17–2.47)	(2.31–2.80)
LAB	4.59 ± 0.21	4.51 ± 0.12	4.23 ± 0.13	4.23 ± 0.12	3.86 ± 0.14	4.55 ± 0.07	4.45 ± 0.11	3.23
	(4.26–5.23)	(4.28–4.74)	(4.04–4.34)	(3.98–4.43)	(3.73–4.05)	(4.48–4.63)	(4.23–4.61)
CLM	3.80 ± 0.07	4.13 ± 0.12	3.78 ± 0.05	3.85 ± 0.18	3.16 ± 0.09	4.10 ± 0.09	4.17 ± 0.14	2.60
	(3.66–3.93)	(3.97–4.51)	(3.74–3.85)	(3.40–4.17)	(3.02–3.25)	(3.97–4.18)	(3.99–4.52)
WM1	1.20 ± 0.04	1.24 ± 0.04	1.13 ± 0.01	1.18 ± 0.05	0.95 ± 0.09	1.24 ± 0.04	1.28 ± 0.06	0.82
	(1.12–1.29)	(1.14–1.30)	(1.12–1.13)	(1.06–1.28)	(0.86–1.06)	(1.19–1.28)	(1.17–1.34)
TOT	230.2 ± 12.1	245.2 ± 7.1	234.3 ± 6.6	227.3 ± 12.5	204.4 ± 7.6	222.4 ± 13.4	233.3 ± 12.6	132
	(201–260)	(233–264)	(230–244)	(205–262)	(193–212)	(205–254)	(207–255)
	50	32	4	50	5	16	21	1
HB	92.6 ± 5.6	101.7 ± 3.8	97.8 ± 1.9	91.3 ± 5.6	83.2 ± 4.4	87.2 ± 6.4	94.8 ± 6.8	56
	(80–103)	(93–109)	(95–99)	(77–104)	(76–87)	(78–101)	(82–106)
	50	32	4	50	5	16	21	1
TAIL	137.6 ± 8.8	145.5 ± 5.1	138.3 ± 6.3	136.0 ± 9.0	121.2 ± 3.3	135.2 ± 9.2	138.5 ± 7.4	76
	(117–158)	(136–159)	(132–147)	(115–161)	(117–125)	(121–153)	(121–150)
	50	32	4	50	5	16	21	1
HF	20.4 ± 0.9	21.1 ± 0.8	20.3 ± 0.5	20.5 ± 1.0	18.0 ± 1.0	20.7* ± 0.5	22.0* ± 0.9	14
	(19–22)	(20–23)	(20–21)	(18–22)	(17–19)	(19.5–21.5)*	(19.5–23)*
	49	32	4	50	5	24	21	1
EAR	19.8 ± 1.0	19.9 ± 0.8	19.0 ± 0.0	18.2 ± 1.0	14.8 ± 0.8	18.0 ± 0.7	18.6 ± 1.0	13
	(17.5–22)	(18–21)	(19)	(17–21)	(14–16)	(16.5–19)	(17–21)
	47	32	4	50	5	24	21	1
Wt	24.9 ± 4.0	27.6 ± 3.1	21.8 ± 1.2	22.0 ± 3.7	13.9 ± 1.4	20.4 ± 4.0	NA	5.8
	(17–39)	(22–34.5)	(20.5–23)	(15–29.5)	(11.5–15)	(16–30.5)
	48	31	4	50	5	15		1
TAIL/HB	1.5 ± 0.1	1.4 ± 0.0	1.4 ± 0.1	1.5 ± 0.1	1.4 ± 0.1	1.6 ± 0.1	1.5 ± 0.1	1.4
	(1.2–1.7)	(1.4–1.5)	(1.3–1.5)	(1.2–1.7)	(1.4–1.5)	(1.4–1.8)	(1.3–1.7)
	50	32	4	50	5	16	21	1