Novel splitting / lumping index reflects the history of species concepts applied to bumblebees (Insecta: Apidae) Author Williams, Paul H text Zoological Journal of the Linnean Society 2022 2022-10-01 196 2 704 719 https://academic.oup.com/zoolinnean/article/196/2/704/6540035 journal article 10.1093/zoolinnean/zlab123 0024-4082 7184415

Table 4. As for Table 1 but for the subgenus Melanobombus

Source	Smith, 1854	von Dalla	Skorikov, 1923	Reinig, 1935	Williams, 1998	Williams et al.,	Combined list of
		Torre, 1896				2020 (‘present’)	candidate species
Species concept	typological	typological/	typological/	polytypic/	polytypic/	evolutionarily inde-
		polytypic	polytypic	interbreeding interbreeding		pendent lineages
Character set	colour	colour/	colour/	morphology	morphology/mate-	molecular/morph-
		morphology morphology			searching behaviour ology/CLGS
Species	rufipes	rufipes	rufipes	[x]	rufipes	rufipes	rufipes
	eximius	eximius	eximius		eximius	eximius	eximius
			latissimus	latissimus			latissimus
		festivus	festivus	[x]	festivus	festivus	festivus
			atocinctus				atocinctus
			terminalis				terminalis
			molleri				molleri
	similis	simillimus	simillimus	[x]	simillimus	simillimus	simillimus
			tonsus				tonsus
			miniatus	miniatus	miniatus	miniatus	miniatus
			flavothoracicus				flavothoracicus
					x	eurythorax	eurythorax
	rufofasciatus rufofasciatus rufofasciatus			rufofasciatus	rufofasciatus	rufofasciatus	rufofasciatus
		prshewalskii	przewalskii	x	x	prshewalskyi	prshewalskyi
					richardsiellus	richardsiellus	richardsiellus
				[x]	friseanus	friseanus	friseanus
		pyrrhosoma	pyrrhosoma	pyrrhosoma	pyrosoma	pyrosoma	pyrosoma
				[x]	formosellus	formosellus	formosellus
		tanguticus	tanguticus	tanguticus	tanguticus	tanguticus	tanguticus
	eriophorus	eriophorus	caucasicus	x	x	eriophorus	eriophorus
		caucasicus					caucasicus
	lapidarius	lapidarius	lapidarius	lapidarius	lapidarius	lapidarius	lapidarius
			semenovianus	semenovianus	semenovianus	semenovianus	semenovianus
		x	incertus	incertus	incertus	incertus	incertus
		alticola	sicheli	sicheli	sichelii	sichelii	sichelii
			alticola				alticola
			cazurroi				cazurroi
			verticiflavus				verticiflavus
			bianchii	[x]	ladakhensis	ladakhensis	ladakhensis
			variopictus				phariensis
			*	*	*	tibeticus	tibeticus
			alagesianus	x	x	alagesianus	alagesianus
			incertoides	x	x	incertoides	incertoides