Two new Pandanus frogs (Guibemantis: Mantellidae: Anura) from northern Madagascar Author Lehtinen, Richard M. 9FCBFB1E-4A52-4747-A212-D6B9513453AC The College of Wooster, Department of Biology, 931 College Mall, Wooster, Ohio, 44691 USA. rlehtinen@wooster.edu Author Glaw, Frank 764DC666-B65D-40C1-BE19-0F5A7294E452 Zoologische Staatssammlung München (ZSM-SNSB), Münchhausenstr. 21, 81247 München, Germany. glaw@snsb.de Author Vences, Miguel 23B8FF04-06AA-43C8-A459-E0FB3BB289F6 Zoological Institute, Technical University of Braunschweig, Mendelssohnstr. 4, 38106 Braunschweig, Germany. m.vences@tu-braunschweig.de Author Rakotoarison, Andolalao A354F4A8-FDD9-4072-9C27-DC17B6ED3664 Mention Zoologie et Biodiversité Animale, Université d’Antananarivo, BP 566, Antananarivo 101, Madagascar. andomailaka@gmail.com Author Scherz, Mark D. DDE9BA71-9261-483D-AC69-6985F388DD04 Zoologische Staatssammlung München (ZSM-SNSB), Münchhausenstr. 21, 81247 München, Germany. & Zoological Institute, Technical University of Braunschweig, Mendelssohnstr. 4, 38106 Braunschweig, Germany. mark.scherz@gmail.com text European Journal of Taxonomy 2018 2018-07-19 451 1 20 journal article 22313 10.5852/ejt.2018.451 aa402c82-b500-429a-966f-f297120019b6 2118-9773 3814542 40E9A16C-69E9-47FA-8588-A9F2FFFB63D9

Table 1 (continued on next page). Morphometric data (in mm) of available specimens of the two new species, Guibemantis albomaculatus sp. nov. (n = 15), and G. woosteri sp. nov. (n = 14), as well as comparative specimens of G. albolineatus (Blommers-Schlösser & Blanc, 1991) (n = 7). HT = holotype, PT = paratype, LT = lectotype, PLT = paralectotype, SD = standard deviation; see Material and methods for other abbreviations.

Catalogue Number	Locality	Status	Sex	SVL	FL	TL	FOL	HL	HW	HAL	HAL/SVL	LLA	TD	ED	FGW	FGL	W3FD	W4TD	IOD	NSD	END	IND
G. albomaculatus sp. nov.
UMMZ 212592 UMMZ 212593 UMMZ 212594 UMMZ 212595 UMMZ 212596 ZSM 0816/2003 / FGMV 2002.735 UMMZ 212585 UMMZ 212587 UMMZ 212588 UMMZ 212591 ZSM 496/2000 / FGMV 2000.356 ZSM 0874/2003 / FGMV 2002.869 ZSM 0893/2003 / FGMV 2002.901 ZSM 0894/2003 / FGMV 2002.904 ZSM 0895/2003 / FGMV 2002.905	Manongarivo Manongarivo Manongarivo Manongarivo Manongarivo Manongarivo Montagne d’Ambre Montagne d’Ambre Montagne d’Ambre Montagne d’Ambre Montagne d’Ambre Montagne d’Ambre Montagne d’Ambre Montagne d’Ambre Montagne d’Ambre	PT PT PT PT PT PT PT PT PT PT PT PT PT PT HT	♀ ♀ ♀ ♂ ♂ ♂ ♂ ♀ ♂ ♀ ♀ ♀ ♀ ♀ ♂	24.1 24.5 25.3 21.6 18.9 22.4 22.9 24.3 20.0 26.7 22.2 24.4 23.3 20.9 22.0	11.0 10.4 10.6 9.4 8.7 10.2 10.1 11.1 9.2 11.3 9.5 10.8 9.4 9.2 9.0	12.1 11.3 12.4 10.9 9.9 11.1 10.5 12.0 9.5 12.8 10.4 11.7 10.6 10.2 10.6	10.0 9.3 10.3 9.1 8.3 9.3 9.3 10.5 8.0 11.0 9.4 10.7 10.1 8.8 8.8	8.1 8.3 8.7 7.7 7.3 7.8 7.7 9.2 6.7 9.7 6.7 8.0 7.8 7.9 7.1	8.2 8.4 8.8 6.6 6.3 7.5 7.9 8.4 6.9 9.3 7.9 8.9 7.7 7.1 7.6	6.1 6.9 6.5 6.1 5.7 5.7 6.0 6.4 4.7 6.6 5.5 6.5 6.5 4.6 5.6	0.3 0.3 0.3 0.3 0.3 0.3 0.3 0.3 0.2 0.2 0.2 0.3 0.3 0.2 0.3	4.7 5.1 4.4 4.5 3.9 5.2 5.7 4.7 3.6 5.1 4.7 5.6 4.9 4.4 5.1	1.4 1.6 1.5 1.5 1.2 1.3 1.5 1.3 1.2 1.2 1.2 1.2 1.1 1.0 1.1	2.9 2.9 3.1 2.8 2.5 2.9 2.8 3.1 2.5 2.9 2.8 3.6 2.8 2.8 2.6	– – – 1.0 1.4 – 1.3 – 1.5 – – – – – 1.4	– – – 5.1 4.1 – 4.9 – 3.7 – – – – – 4.4	0.8 0.9 1.1 0.7 1.0 1.2 0.9 0.9 0.9 0.7 1.1 1.4 1.2 0.8 1.2	0.6 0.9 0.8 0.6 0.8 1.0 0.8 0.6 0.6 0.5 0.6 0.9 0.8 0.6 0.7	2.9 3.5 3.2 2.6 2.9 3.3 3.4 3.2 2.9 3.1 3.0 3.7 3.6 2.5 2.9	0.6 0.8 0.7 0.6 0.9 0.9 0.7 0.7 0.9 0.5 0.9 1.0 0.8 0.9 0.8	3.0 2.8 2.7 2.2 2.4 2.7 2.7 2.9 2.4 2.9 2.5 2.7 2.4 2.3 2.3	2.4 1.9 2.0 2.2 1.9 2.3 2.2 1.5 1.8 2.2 2.3 2.5 2.4 2.4 2.4
G. woosteri sp. nov.
UMMZ 212601 UMMZ 212602 UMMZ 212604 UMMZ 212605 UMMZ 212606 UMMZ 212613 UMMZ 212615 UMMZ 212617 UMMZ 191339 UMMZ 191340 UMMZ 191343	Marojejy Marojejy Marojejy Marojejy Marojejy Marojejy Marojejy Marojejy Ambatovaky Reserve Ambatovaky Reserve Ambatovaky Reserve	PT PT PT PT PT PT PT PT – – –	♂ ♂ ♂ ♂ ♂ ♀ ♀ ♀ ♂ ♀ ♂	21.8 21.5 21.8 21.1 20.4 23.9 23.0 22.5 21.7 24.5 21.5	10.2 9.9 9.8 9.5 9.6 10.5 10.4 10.1 10.4 10.8 9.5	11.4 11.2 11.0 10.5 11.0 11.6 11.7 11.5 11.5 12.1 10.2	9.9 9.9 10.1 9.5 9.2 9.0 10.5 10.2 8.6 10.1 8.1	7.9 7.5 8.5 7.0 7.7 8.0 8.0 8.2 8.3 9.0 7.7	7.0 6.9 7.3 7.4 7.0 7.5 8.0 7.0 6.8 7.1 6.7	6.1 6.1 5.7 6.1 5.7 6.3 6.6 5.7 5.3 6.2 5.2	0.3 0.3 0.3 0.3 0.3 0.3 0.3 0.3 0.2 0.3 0.2	5.1 4.4 4.6 4.7 4.7 4.5 5.2 5.1 4.4 4.3 3.7	1.3 1.3 1.4 1.2 1.5 1.4 1.5 1.6 1.2 1.3 1.3	3.2 2.8 3.2 2.8 2.9 3.0 3.3 2.6 2.8 2.9 3.0	1.4 1.3 1.5 1.7 1.4 – – – 1.7 – 1.2	4.8 4.4 4.7 5.1 4.1 – – – 4.2 – 4.1	0.9 0.9 0.9 1.0 1.1 1.0 1.0 0.8 0.8 0.9 0.7	0.7 0.8 0.6 0.8 0.7 0.8 0.9 0.8 0.7 0.6 0.6	2.9 2.9 2.6 2.8 3.2 3.1 2.8 3.1 2.8 3.1 2.8	1.0 0.7 0.8 0.8 0.7 0.7 1.0 0.7 0.9 1.2 0.8	2.5 2.6 2.2 2.1 2.2 2.3 2.2 2.4 2.6 2.9 2.8	2.0 2.4 1.9 2.2 2.1 2.2 2.7 2.2 2.2 2.3 1.9
UMMZ 191344 ZSM 5062/2005 / ZCMV 2011 ZSM 5063/2005 / ZCMV 2044	Ambatovaky Reserve Marojejy Marojejy	– PT HT	♂ ♂ ♀	19.4 20.8 24.9	9.2 10.3 10.5	10.4 10.6 11.5	9.2 9.0 10.1	7.6 6.9 8.8	6.0 7.3 8.3	5.4 6.0 5.8	0.3 0.3 0.2	4.8 5.1 5.4	1.1 1.1 1.3	2.9 3.1 3.4	1.1 1.6 –	3.7 5.0 –	1.0 1.1 1.2	0.6 0.8 1.1	2.8 2.2 3.3	0.7 0.7 1.1	1.8 2.3 2.7	2.0 2.3 2.6
G. albolineatus (Blommers-Schlösser & Blanc, 1991)
ZSM 171/2005 / FGZC 2432 ZSM 172/2005 / FGZC 2433 MNHN 1972.2074 MNHN 1972.2082 MNHN 1972.2077 MNHN 1972.2079 MNHN 1972.2080	Andohahela Andohahela Andohahela Andohahela Andohahela Andohahela Andohahela	– – PT PT PT PT PT	♂ ♂ ♂ ♀ ♀ juv. ♀	21.3 20.9 19.8 19.0 22.2 16.2 18.9	9.5 10.0 9.1 8.0 9.3 7.8 9.1	10.9 10.7 9.8 9.8 10.8 8.9 9.5	9.7 9.9 7.4 8.9 9.0 7.9 8.2	6.4 6.6 6.7 7.4 7.5 5.8 5.7	7.6 7.4 6.8 6.9 7.2 6.5 6.4	7.2 7.1 5.5 5.5 6.2 5.2 5.6	0.34 0.34 0.28 0.29 0.28 0.32 0.30	5.1 5.1 4.3 4.7 4.9 4.5 4.4	1.2 1.0 1.1 1.1 1.0 1.0 1.0	2.8 2.8 2.5 2.6 2.9 2.9 2.5	1.8 2.1 1.4 – – – –	5.0 5.7 4.7 – – – –	1.3 1.0 1.0 0.9 1.2 0.9 1.0	1.2 0.9 0.9 0.8 0.8 0.5 0.7	3.2 2.9 2.4 2.8 3 2.5 2.5	1.2 1.3 0.8 0.6 0.6 0.7 0.8	2.2 2.1 2.1 2.1 2.2 1.9 2.0	2.4 2.6 2.3 2.3 2.7 2.3 2.4