| Achene surface | polygons surrounded by papillae | irregularly shallow reticulations |
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| Leaf epidermis | upper leaf epidermis: epidermal cells polygonal, | upper leaf epidermis: epidermal cells irregular, |
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| stomatal apparatus paracytic lower leaf epidermis: | s t o m a t a l a p p a r a t u s a n i s o c y t i c l o w e r l e a f |
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| stomata size 18.6 (17.8–19.7) µm × 15.7 (14.5–16.9) | epidermis: stomata size 25.4 (23.1–26.8) µm × |
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| µm, stomata density 253 number/mm 2, guard cells | 19.6 (18.2–20.9) µm, stomata density 197 |
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| enclosed by 2 subsidiary cells mostly unequal in size | number/mm 2, guard cells enclosed by 2 equal |
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| or sometimes 3 obviously unequal in size | subsidiary cells |
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| Pollen grains | L-morph: lumen polygonal, uniform, 7.64 ± 0.351 | L-morph: lumen irregular, unequal, 8.42 ± 0.719 |
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| µm in size, 7 (6–8) lumina across the diameter, | µm, in size, 9 (8–10) lumina across the diameter, |
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| columellae in a lumen 19 (16–27) S-morph: lumen | columellae in a lumen 24 (15–31) S-morph: |
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| polygonal, uniform, 10.21 ± 0.512 µm in size, 8 (7– | lumen polygonal, unequal, 11.63 ± 0.823 µm in |
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| 9) lumina across the diameter, columellae in a lumen | s i z e, 5 (6–7) l u m i n a a c r o s s t h e d i a m e t e r, |
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| 37 (26–48) | columellae in a lumen asymmetrical, 33 (24–45) |
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| Tepal inner surface | epidermal cells 45–72 µm × 14–25 µm, longitudinal | e p i d e r m a l c e l l s 3 5 – 5 5 µ m × 8– 1 5 µ m, |
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| direction with striate cuticles, anticlinal wall straight | longitudinal direction with striate cuticles, |
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| or sinuolate, surface granular | anticlinal wall sinuate, surface with intensely |
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| | developed ridge |
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