The Lutrinae (Mustelidae, Carnivora, Mammalia) from the Upper Miocene to the Lower Pleistocene deposits of Pakistan Author Mahmood, Khalid Dr. Abu Bakr Fossil Display & Research Centre, Department of Zoology, University of the Punjab, Quid-e-Azam Campus, Lahore 54590 (Pakistan) and Department of Zoology, Govt. Islamia Graduate College Kasur, Punjab (Pakistan) khalidkasuri1@gmail.com Author Morlo, Michael Forschungsinstitut Senckenberg, Abt. Messelforschung und Mammalogie, Senckenberganlage 25, 60325 Frankfurt (Germany) michael.morlo@senckenberg.de Author Abbas, Sayyed Ghyour Department of Zoology, University of Sialkot, Sialkot, Punjab (Pakistan) sayyed.ghyour@uskt.edu.pk & ghyour.kazmi@gmail.com Author Babar, Muhammad Adeeb Department of Zoology, University of Okara, Okara, Punjab (Pakistan) babar.441@googlemail.com Author Khan, Muhammad Akbar Dr. Abu Bakr Fossil Display & Research Centre, Department of Zoology, University of the Punjab, Quid-e-Azam Campus, Lahore 54590 (Pakistan) akbar.zool@pu.edu.pk text Geodiversitas 2025 2025-03-06 47 5 301 311 https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/geodiversitas2025v47a5.pdf journal article 10.5252/geodiversitas2025v47a5 1df16561-1d43-465c-b811-d2da4b852f2d 1638-9395 15019534 urn:lsid:zoobank.org:pub:EEED70E5-D9C1-4642-B363-E5D22CDBB19C

TABLE 2. — Comparative measurements of Enhydriodon cf. falconeri, Sivaonyx sarwari n. sp. and Sivaonyx bathygnathus (Lydekker, 1884) (in mm). Comparative data taken from Pilgrim (1932), Colbert (1935), Chow (1961), Qi (1983), Zong (1997), Werdelin (2003), Morales & Pickford (2005), Morales et al. (2005), Pickford (2007), Peigné et al. (2008), Geraads et al. (2011), Werdelin & Manthi (2012), Grohé et al. (2013, 2022), Werdelin & Lewis (2013), Peigné (2016), Koufos et al. (2018) and Valenciano & Govender (2020). *, studied specimens; **, measurements as preserved; est., estimated measurements.

Taxa	SpecimenNo	Nature	Length	Width	RatioW/L
Enhydriodon cf. falconeri	PUPC 83/01*	rm1	18.0	11.0	0.69
	NHM M 15397	rm1	18.6	10.9	0.59
	Chow, 1961	lm1	20.0	13.0	0.65
E. sivalensis Falconer, 1868	IPSMG 1949.187	rm1	22.0**	13.0**	0.59
	GSI D 161	rm1	21.60	13.1	0.61
Enhydriodon omoensis Grohé, Uno &	L 56-1	m1	32.1	18.3	0.57
Boisserie, 2022	OMO 18-1972-99	m1	31.7**	19.1**	0.60
Enhydriodon dikikae Geraads, Alemseged,	DIK-56-9 (holotype)	m1	30.0**	20.0**	0.66
Bobe & Reed, 2011	DIK-24-15	m1	26.0	16.2	0.62
Enhydriodon afman Werdelin & Lewis, 2013 KNM-ER 3110 (holotype)		m1	26.4	16.5	0.63
Enhydriodon sp.	OMO 3/0 10084	m1	26.4**	11.8**	0.45
Sivaonyx sarwari n. sp.	PUPC 17/64*	lP4	12.50	10.00	0.80
Sivaonyx bathygnathus (Lydekker, 1884)	GSI K13.14	lP4	14.00 est.	13.40	0.96
	GSI D 157	lP4	14.50	13.4	0.92
	IVPP V 6886.3	rP4	13.0	13.8	1.06
	IVPP V 6886.4	lP4	13.2	13.5	1.02
S. gandakasensis Pickford, 2007	GSP 4616	rP4	13.5	11.5	0.85
Sivaonyx hessicus Lydekker, 1890	PV-9002	lP4	12.50	10.7	0.85
S. soriae Morales & Pickford, 2005	BAR 1720’00	P4	14.8	15.0	1.01
S. hendeyi Morales, Pickford & Soria, 2005	SAM-PQL 50000B (holotype)	P4	16.9	17.4	1.03
S. ekecaman (Werdelin, 2003)	KNM-KP 10034A (holotype)	P4	16.5	–	–
	BAR 566’05	P4	14.7	17.5	1.19
E. falconeri Pilgrim, 1932	NHM M 4847	lP4	14.9	15.8	1.06
E. sivalensis	GSI NRV 2/468	rP4	17.3	18.6	1.08
	NHM M 37153	rP4	16.7 est.	17.2	1.03
	NHM M 37154	rP4	17.6	18.9	1.07
	NHM M 37154	lP4	17.6	18.9	1.07
	NHM M 37155	lP4	18.2	18.5	1.01
	GSI RCS 777A cast	lP4	16.5	18.4	1.11
	GSI RCS 777A cast	rP4	17.6	18.6	1.05
E. omoensis	P 791-18 (holotype)	P4	25.8**	26.7**	1.03
E. dikikae Geraads, Alemseged, Bobe &	DIK-56-9 (holotype)	P4	20.5	22.7	1.11
Reed, 2011
E. cf. dikikae	KNM-KP 49887	P4	18.4	19.5	1.06
Sivaonyx bathygnathus (Lydekker, 1884)	PUPC 16/29*	lp4	10.4	6.70	0.64
S. bathygnathus	PUPC 2003/15	p4	7.50	5.50	0.73
	GSI D 156	lp4	9.6	5.80	0.60
	GSI D 244	rp4	10.1	6.40	0.63
	IVPP V 6886.3	rp4	13.0	13.8	1.06
	IVPP V 6886.4	lp4	13.2	13.5	1.02
	IVPP V 6886.8+13	lp4	8.80	6.00	0.68
	IVPP V 6886.9	lp4	8.40	6.00	0.71
	IVPP V 6886.10	lp4	8.80	6.50	0.74
	PDYV 1585	lp4	8.70	5.00	0.57
S. hendeyi Morales, Pickford & Soria, 2005	SAM-PQL 50000A (holotype)	p4	12.0	9.40	0.78
	SAM-PQL-9138	p4	13.8	9.90	0.72
S. ekecaman	BAR 720’03	p4	11.1	8.40	0.76
S. africanus Stromer, 1931	BSPG 1930 XI 1 (holotype)	p4	12.0	9.80	0.82