Description of three new species previously identified as Stolephorus bengalensis (Dutt & Babu Rao, 1959) or Stolephorus insularis Hardenberg, 1933 and a re-description of S. bengalensis (Chordata, Osteichthyes, Clupeiformes, Engraulidae) Author Hata, Harutaka https://orcid.org/0000-0002-6996-3628 National Museum of Natural History, Smithsonian Institution, 10 k2795502@kadai.jp Author Lavoue, Sebastien https://orcid.org/0000-0003-4798-6666 th Author Motomura, Hiroyuki https://orcid.org/0000-0002-7448-2482 and Constitution Ave, NW Washington, DC 20560, USA text ZooKeys 2022 2022-09-15 1121 145 173 http://dx.doi.org/10.3897/zookeys.1121.84171 journal article http://dx.doi.org/10.3897/zookeys.1121.84171 1313-2970-1121-145 7AA4A7A27F825B08A29992271115479D

Table 2. Meristics of specimens of Stolephorus bengalensis and Stolephorus eldorado sp. nov.

	Stolephorus bengalensis			Stolephorus eldorado sp. nov.
	Neotype of Anchoviella baganensis bengalensis	Non-types		Holotype	Paratypes
	USNM 276476	n = 46		KAUM-I. 94517	n = 57
*one specimen with 4 unbranched anal-fin rays.
Standard length (mm)	45.8	30.8-58.7	Modes +/- SD	44.4	37.5-58.8	Modes +/- SD
Dorsal-fin rays (unbranched)	3	3	3 +/- 0	3	3	3 +/- 0
Dorsal-fin rays (branched)	12	11-14	13 +/- 0.7	13	11-14	13 +/- 0.6
Anal-fin rays (unbranched)	3	3	3 +/- 0	3	3*	3 +/- 0.1
Anal-fin rays (branched)	18	16-20	18 +/- 0.9	18	16-19	18 +/- 0.6
Pectoral-fin rays (unbranched)	1	1	1 +/- 0	1	1	1 +/- 0
Pectoral-fin rays (branched)	11	10-12	11 +/- 0.7	12	9-13	11 +/- 0.8
Pelvic-fin rays (unbranched)	1	1	1 +/- 0	1	1	1 +/- 0
Pelvic-fin rays (branched)	6	6	6 +/- 0	6	6	6 +/- 0
Gill rakers on 1st gill arch (upper)	16	17-19	18 +/- 0.8	18	16-21	18 +/- 1.1
Gill rakers on 1st gill arch (lower)	24	23-27	25 +/- 1.1	26	23-28	25 +/- 1.0
Gill rakers on 1st gill arch (total)	40	40-45	44 +/- 1.5	44	40-47	42 +/- 1.7
Gill rakers on 2nd gill arch (upper)	13	11-15	13 +/- 0.8	12	10-14	13 +/- 0.8
Gill rakers on 2nd gill arch (lower)	22	21-25	23 +/- 0.9	24	20-24	23 +/- 1.0
Gill rakers on 2nd gill arch (total)	35	33-39	36 +/- 1.5	36	30-38	36 +/- 1.5
Gill rakers on 3rd gill arch (upper)	10	10-12	11 +/- 0.6	10	8-12	10 +/- 0.8
Gill rakers on 3rd gill arch (lower)	13	13-15	13 +/- 0.6	13	12-14	13 +/- 0.6
Gill rakers on 3rd gill arch (total)	23	23-27	24 +/- 1.1	23	20-26	23 +/- 1.1
Gill rakers on 4th gill arch (upper)	9	7-9	8 +/- 0.6	8	7-10	8 +/- 0.7
Gill rakers on 4th gill arch (lower)	10	10-12	11 +/- 0.5	11	9-12	11 +/- 0.8
Gill rakers on 4th gill arch (total)	19	17-21	19 +/- 1.0	19	16-22	18 +/- 1.3
Gill rakers on posterior face of 3rd gill arch	6	4-7	5 +/- 0.7	5	4-7	5 +/- 0.7
Prepelvic scutes	7	5-7	6 +/- 0.5	6	5-7	6 +/- 0.6
Scale rows in longitudinal series	35	34-36	35 +/- 0.7	34	34-36	35 +/- 0.7
Transverse scales	8	8	8 +/- 0	8	8-9	8 +/- 0.3
Pseudobranchial filaments	broken	13-18	16 +/- 1.3	17	14-18	16 +/- 1.2
Total vertebrae	40	40-41	40 +/- 0.4	39	38-40	39 +/- 0.7