Redescriptions of Pugettia quadridens (De Haan, 1837) and P. intermedia Sakai, 1938 (Crustacea: Brachyura: Epialtidae) with description of a new species Author Ohtsuchi, Naoya Author Kawamura, Tomohiko text Zootaxa 2019 2019-09-20 4672 1 1 68 journal article 25435 10.11646/zootaxa.4672.1.1 24c8554d-e477-4440-8e3f-17af20c1ddc8 1175-5326 3451561 01B7FFA9-1EC8-41A7-8DDE-418D9F4882B8

TABLE 2. Summary of differences among Pugettia quadridens (De Haan, 1837), P. intermedia Sakai, 1938, P. ferox n. sp., P. pellucens Rathbun, 1932, and P. vulgaris Ohtsuchi, Kawamura & Takeda, 2014.

Characters	P. quadridens	P. intermedia	P. ferox n. sp.	P. pellucens	P. vulgaris
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* not clear in small specimens ** not clear in immature specimens
Protuberances on gastric and	often rudimentary; each	often distinct; each protogastric	often distinct; each	absent	absent
branchial regions*	protogastric protuberance	protuberance more separated	protogastric protuberance
	separated equally from	from mesogastric protuberance	separated equally from
	meso- and metagastric	than from metagastric	meso- and metagastric
	protuberance	protuberance	protuberance
Mesobranchial region*	lower than gastric region,	as high as gastric region, with	as high as gastric region,	moderately elevated,	moderately elevated,
	with 2 rudimentary tubercles	2–3 distinct tubercles	with 2–3 distinct tubercles	unarmed	unarmed
Metabranchial region*	unarmed	with a large tubercle	with a large tubercle	unarmed	unarmed
Cardiac and intestinal regions*	fused	separated	separated	separated	separated
Rows of hooked setae on	often absent in larger males	always present, posterior end	always present	always present	always present
the anterior slope of gastric		of former positioned posterior
region and a group of setae on		to hepatic lobe basis
anterolateral carapace submargin
Relative length of pseudorostral	0.2–0.3	0.2–0.3	0.1–0.2	0.4–0.5	0.3–0.4
spines (PRL/PCL, in full-grown
males)
Supraorbital eave	moderately inclined dorsally;	noticeably inclined dorsally;	moderately inclined	nearly straight, truncate	nearly straight, truncate
	lateral margin distinctly	nearly straight or shallowly	dorsally; lateral margin
	concave, truncate on	concave, distinctly truncated on	variably sinuous
	posterior end	posterior end
Orbital hiatus	small, triangular notch	deep, wide concavity	deep, wide concavity	deep, narrow sulcus	deep, wide concavity
Hepatic lobe	moderately compressed	nearly spiniform in distal	moderately compressed	moderately compressed	moderately compressed
	in entire length; posterior	half; posterior margin bent	in entire length; posterior	in entire length;	in entire length;
	margin weakly convex;	outwards; posterior end right	margin nearly straight;	posterior margin weakly	posterior margin slightly
	posterior end right above	above midpoint of pleural	posterior end right above	sinuous; posterior end	concave; posterior end
	posterior end of pleural	suture	midpoint of posterior end	right above midpoint of	right above midpoint of
	suture		of pleural suture	pleural suture	pleural suture
Relative position of	0.41±0.03	0.33±0.03	0.36±0.02	0.3	0.3
epibranchial spine (ESL/PCL)	(0.35–0.46)	(0.27–0.36)	(0.31–0.38)
Penultimate article of antennal	not carinate on mesial,	carinate on lateral, mesial	carinate on mesial, lateral	arinate on lateral margin	carinate on mesial, lateral
peduncle	lateral margins	margins	margins (mesial often		margins (mesial often
			rudimentary)		rudimentary)
Lateral margin of basal	thickened at distolateral	not thickened at	not thickened at	not thickened at	not thickened at distolateral
antennal article	angle, not markedly	distolateral angle, well	distolateral angle, well	distolateral angle, well	angle, well extended laterally,
	extended laterally,	extended laterally,	extended laterally,	extended laterally, faintly	faintly sinuous, proximal end
	concave, proximal end	sinuous, proximal end	sinuous, proximal end	sinuous, proximal end	separated from suborbital
	fused with suborbital	separated from suborbital	separated from suborbital	separated from suborbital	region
	region	region	region by (often pointed)	region
			expansion
Third maxilliped	ischium with a shallow,	ischium with a	ischium with a shallow,	ischium with a shallow,	ischium with a shallow,
	broad, longitudinal	longitudinal groove;	broad, longitudinal	broad, longitudinal	broad, longitudinal
	depression; expod	expod immediately	depression; expod	depression; expod	depression; expod gradually
	gradually narrowed in	narrowed in distal one-	immediately narrowed in	gradually narrowed in	narrowed in distal half
	distal half	third, compressed in	distal two-fifth	distal half
		lateral half
Cheliped merus armatures on	3 low teeth (distalmost	3–6 large, lamellar teeth	3–6 large teeth	3 lamellar teeth	3 lamellar teeth (distalmost
dorsal and outer longitudinal	lowest) on dorsal ridge;	(distalmost highest) on	(distalmost highest)	(distalmost highest)	highest) on dorsal ridge; 3 or
ridge*	unarmed on outer ridge	dorsal ridge; 4–7 low	on dorsal ridge; 2 or 3	on dorsal ridge; 3 or 4	4 low tubercles on outer ridge
		tubercles on outer ridge	rudimentary teeth on	rudimentary tubercles on
			outer ridge	outer ridge
Cheliped carpus dorsal ridge*	obtuse, unarmed	sharply dentate	obtuse, often tuberculate	bi- or tri-tuberculate	bi- or tri-tuberculate
Chela (full-grown males)	fingers widely gaped in	fingers widely gaped in	fingers narrowly gaped	fingers narrowly gaped	fingers narrowly gaped in
	proximal 2/3; movable	proximal 1/2 or entire	in proximal 1/2 or entire	in entire length; movable	entire length; movable finger
	finger with 2, 3 large	length; movable finger	length; movable finger	finger uniformly dentate	uniformly dentate
	teeth at proximal 1/3	uniformely dentate in	uniformly dentate
		distal 2/3
Ambulatory legs meri	generally smooth,	generally tomentose, upper	generally smooth, upper	generally smooth, upper	generally smooth, upper distal
	upper distal tubercle	distal tubercle distinct	distal tubercle moderate	distal tubercle distinct	tubercle distinct
	faint
Depression on extensor	faint	distinct	distinct	distinct	distinct
surface of ambulatory
legs carpi
Third to fifth ambulatory	subcylindrical	weakly flattened	weakly flattened	weakly flattened	subcylindrical
legs propodi
Spinulation on flexor	large	small, sometimes reduced	small, sometimes	large	large
surface of first and		in large specimens	reduced in large
second ambulatory legs			specimens
dactyli
Lateral margin of male	elevated higher	elevated as high as mesial	elevated as high as	elevated as high as mesial	elevated as high as mesial margin
first gonopod	than mesial margin	margin	mesial margin	margin
	dinstinguishes
Small ridge on basis of	absent	absent	present	absent	absent
male first gonopod dorsal
lobe**
Mesial lobe of male first	short, projecting	variably long, directed	short, directed anteriorly	short, directed anteriorly	short, directed anteriorly obliquely,
gonopod	nearly perpendicular	anteriorly, subparallel or	obliquely, weakly	obliquely, separated from	separated from dorsal lobe by
	to dorsal lobe, curled	crossing with dorsal lobe,	curved downwards,	dorsal lobe by narrow	narrow to wide hiatus
	downwards, separated	separated from dorsal lobe	separated from dorsal	hiatus
	from dorsal lobe by	by narrow hiatus	lobes by variably wide
	wide hiatus		hiatus
Female gonopore	comma-shaped	elongate	comma-shaped	comma-shaped	comma-shaped