A biogeographical puzzle: description of two new frog species (Microhylidae: Callulops and Choerophryne) from boulder-field habitat of Dauan Island, Torres Strait, Australia Author Hoskin, Conrad J. text Zootaxa 2025 2025-09-19 5693 4 501 523 https://doi.org/10.11646/zootaxa.5693.4.3 journal article 312737 10.11646/zootaxa.5693.4.3 b42c132e-98a0-4f4b-8959-8b1398c759ef 1175-5326 17324163 A2682232-0FBA-425D-80A4-B54B722020F2

TABLE 1. Morphological measurements and ratios for the type specimens of Callulops gobakula sp. nov. (QM J98925– 28) and Choerophryne koeypad sp. nov. (QM J98929–32). All measurements are in mm. See Methods for explanation of traits.

	Callulops gobakula sp. nov.				Choerophryne koeypad sp. nov.
	J98925 Holo. Male	J98926 Para. Male	J98927 Para. Female	J98928 Para. Female	J98929 Holo. Male	J98930 Para. Male	J98931 Para. Male	J98932 Para. Male
......continued on the next page
Measures
SVL	51.1	48.5	51.8	50.8	21.0	20.0	21.2	21.1
HW	17.8	17.5	18.2	17.8	7.1	7.7	7.3	7.9
HL	15.9	15.7	16.3	15.6
SN	6.2	5.6	6.2	5.6	2.9	2.8	3.0	2.8
EN	3.3	3.6	4.2	3.6	2.0	1.8	1.9	1.9
IN	4.0	4.0	4.2	4.1	1.7	1.6	1.6	1.6
EYE	5.0	4.5	4.8	4.6	2.2	2.3	2.1	2.3
TYM	3.1	3.3	4.2	3.5
ArmL	22.5	22.0	22.1	22.6	9.8	10.0	9.5	10.2
FA	21.0	20.7	20.6	21.2	4.9	4.9	4.8	5.3
HandL	13.4	12.2	13.4	12.7	4.9	5.1	4.7	4.9
F3L	8.3	7.7	8.1	8.7	2.9	3.1	3.0	2.9
3FD	1.4	1.2	1.5	1.3	1.3	1.4	1.0	1.3
1FD	1.2	1.2	1.3	1.2	0.8	0.8	0.6	0.8
TL–knee	20.5	19.9	21.2	20.5	8.3	8.4	8.9	8.8
TL–fold	18.1	18.2	18.6	19.0
FootL	21.5	20.6	21.0	20.0	6.7	6.7	6.6	6.5
T4L	14.1	12.2	12.7	12.7	3.4	3.8	3.5	3.8
4TD	1.5	1.3	1.6	1.4	1.2	0.9	0.7	0.8
1TD	1.1	1.2	1.1	1.0	1.0	0.9	0.7	0.8
Ratios
HW/SVL	0.35	0.36	0.35	0.35	0.34	0.38	0.35	0.37
HL/SVL	0.31	0.32	0.31	0.31
HL/HW	0.90	0.90	0.90	0.88
SN/SVL	0.12	0.12	0.12	0.11	0.14	0.14	0.14	0.13
EN/SVL	0.064	0.073	0.080	0.071	0.096	0.092	0.090	0.091
IN/SVL	0.078	0.082	0.081	0.081	0.080	0.081	0.074	0.077
EN/IN	0.82	0.90	1.00	0.87	1.20	1.13	1.22	1.17
EYE/SVL	0.097	0.093	0.093	0.091	0.106	0.117	0.101	0.107
TYM/SVL	0.061	0.067	0.082	0.068
ArmL/SVL	0.44	0.45	0.43	0.44	0.46	0.50	0.45	0.48
FA/SVL	0.18	0.20	0.17	0.19	0.23	0.24	0.23	0.25
HandL/ SVL	0.26	0.25	0.26	0.25	0.23	0.25	0.22	0.23
F3L/SVL	0.16	0.16	0.16	0.17	0.14	0.15	0.14	0.14
3FD/SVL	0.027	0.025	0.028	0.025	0.063	0.070	0.049	0.060
1FD/SVL	0.023	0.024	0.024	0.024	0.039	0.038	0.028	0.036
TL–knee/ SVL	0.40	0.41	0.41	0.40	0.39	0.42	0.42	0.42
TL–fold/ SVL	0.35	0.38	0.36	0.37
FootL/SVL	0.42	0.42	0.41	0.39	0.32	0.33	0.31	0.31
T4L/SVL	0.28	0.25	0.24	0.25	0.16	0.19	0.17	0.18
4TD/SVL	0.029	0.026	0.031	0.028	0.058	0.046	0.034	0.040
1TD/SVL	0.022	0.025	0.022	0.020	0.047	0.043	0.033	0.037
3FD/4TD	0.92	0.95	0.92	0.90	1.08	1.52	1.42	1.51