Diagnostic male secondary sexual structures and wing patterns in the Thereus genena species group (Lepidoptera: Lycaenidae: Theclinae) with the description of nine new species from Colombia, Peru, Brazil and French Guiana Author Faynel, Christophe 0000-0001-9330-4633 16, rue des Aspres, 34160 Montaud, France. christophe.faynel@wanadoo.fr Author Huertas, Blanca 0000-0001-9330-4633 Natural History Museum, Cromwell Road, London SW 75 BD, UK. b.huertas@nhm.ac.uk Author Crom, Jean-François Le 0000-0001-9448-9019 calle 61 # 37 - 31, 111321 Bogotá, Colombia. jflecrom@hotmail.com Author O’Brien, Richard 0009-0008-2651-2502 Natural History Museum, Cromwell Road, London SW 75 BD, UK. richard.obrien1@nhm.ac.uk Author Fåhraeus, Christer 0000-0002-2990-3161 Fåhraeus Institute AB, Stortorget 1, 222 23 Lund. & Biodiversity, Department of Biology, Lund University, Lund, Sweden. christer.fahraeus@eqlpharma.com text Zootaxa 2025 2025-12-10 5728 2 201 246 http://dx.doi.org/10.11646/zootaxa.5665.2.10 journal article 313937 10.11646/zootaxa.5728.2.1 ece12af6-cf96-43cf-86d6-27c832341bf1 1175-5326 17894360 2E6171E0-E7A7-430E-BCC0-2C583209A94F

TABLE 3. Comparison of intra- and interspecific p-distances in percent (average ± SD) for the genena group. The alignment is the same as we used for the Iqtree runs. Number of specimens examined within parenthesis. Species ranking by same order as in the text.

	T. endera (12)	T. geminus (2)	T. cacao (2)	T. antecum (3)	T. genena (4)	T. tiasa (5)	T. angulus (4)	T. borbaensis (2)	T. chontachaca (4)	T. ramirezi (2)	T. ortaloides (4)	T. confusus (6)	T. praxioides (8)	T. praxis (3)	T. ortalus (6)	T . aguacatal (6)
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T. endera (12)	1.08 ± 0.53
T. geminus (2)	3.47 ± 0.34	0.15 ± 0.00
T. cacao (2)	2.24 ± 0.22	2.99 ± 0.10	0.00 ± 0.00
T. antecum (3)	6.56 ± 0.36	6.83 ± 0.25	5.21 ± 0.23	0.31 ± 0.27
T. genena (4)	4.44 ± 0.15	4.21 ± 0.10	3.82 ± 0.00	6.74 ± 0.22	0.00 ± 0.00
T. tiasa (5)	4.99 ± 0.26	4.42 ± 0.20	4.19 ± 0.08	7.20 ± 0.24	2.75 ± 0.26	0.85 ± 0.46
T. angulus (4)	4.49 ± 0.31	3.90 ± 0.37	3.90 ± 0.14	6.82 ± 0.25	2.98 ± 0.35	2.97 ± 0.35	1.02 ± 0.53
T. borbaensis (2)	4.14 ± 0.16	4.36 ± 0.11	3.59 ± 0.09	6.81 ± 0.24	4.64 ± 0.08	4.56 ± 0.11	4.25 ± 0.22	0.30 ± 0.00
T. chontachaca (4)	4.82 ± 0.19	4.53 ± 0.08	4.45 ± 0.00	7.12 ± 0.08	5.07 ± 0.00	5.31 ± 0.23	4.93 ± 0.36	4.22 ± 0.08	0.00 ± 0.00
T. ramirezi (2)	4.47 ± 0.14	4.52 ± 0.11	4.13 ± 0.00	7.55 ± 0.07	4.79 ± 0.17	5.34 ± 0.17	4.74 ± 0.29	4.96 ± 0.09	2.77 ± 0.02	0.31 ± 0.00
T. ortaloides (4)	6.34 ± 0.69	5.71 ± 0.34	5.66 ± 0.80	8.78 ± 0.73	5.93 ± 0.74	6.66 ± 0.27	5.50 ± 0.35	6.18 ± 0.82	6.14 ± 0.25	6.69 ± 0.32	0.16 ± 0.17
T. confusus (6)	6.97 ± 0.33	6.56 ± 0.15	6.19 ± 0.08	8.96 ± 0.23	6.68 ± 0.07	6.74 ± 0.12	6.20 ± 0.22	7.26 ± 0.15	6.81 ± 0.12	7.01 ± 0.25	2.66 ± 0.45	0.54 ± 0.57
T. praxioides (8)	6.58 ± 0.38	5.99 ± 0.37	6.31 ± 0.30	9.07 ± 0.36	6.54 ± 0.24	6.22 ± 0.29	5.68 ± 0.48	6.47 ± 0.42	6.13 ± 0.42	6.83 ± 0.20	4.36 ± 0.74	6.09 ± 0.37	1.19 ± 0.64
T. praxis (3)	8.27 ± 0.37	7.18 ± 0.12	7.50 ± 0.14	10.07 ± 0.23	7.84 ± 0.11	8.41 ± 0.33	7.69 ± 0.56	8.63 ± 0.16	8.31 ± 0.02	8.42 ± 0.32	5.08 ± 0.70	6.70 ± 0.29	5.31 ± 0.49	0.61 ± 0.27
T. ortalus (6)	7.19 ± 0.30	6.24 ± 0.18	6.55 ± 0.12	9.21 ± 0.24	6.54 ± 0.36	7.02 ± 0.33	6.14 ± 0.17	6.46 ± 0.14	6.62 ± 0.14	6.34 ± 0.26	3.22 ± 1.12	5.60 ± 0.34	5.11 ± 0.40	5.84 ± 0.30	0.42 ± 0.34
T. aguacatal (6)	7.42 ± 0.30	6.55 ± 0.15	6.87 ± 0.18	10.13 ± 0.16	7.34 ± 0.34	7.66 ± 0.36	6.83 ± 0.38	7.50 ± 0.22	7.71 ± 0.24	7.63 ± 0.38	4.31 ± 0.43	5.50 ± 0.60	4.81 ± 0.39	6.06 ± 0.42	4.94 ± 0.43	0.29 ± 0.56