Three new species and a new record of pheretimoid earthworms (Crassiclitellata, Megascolecidae) from Misamis Oriental, Philippines, with data from the mitochondrial genome Author Aspe, Nonillon M. 0000-0003-0025-6017 College of Fisheries and Marine Sciences, Mindanao State University at Naawan, Naawan 9023, Misamis Oriental, Philippines Author Zhao, Huifeng 0000-0003-4243-9671 Hebei Key Laboratory of Animal Diversity, College of Life Science, Langfang Normal University, Langfang 065000, China & Key Laboratory of Wetland Ecology and Environment, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, China Author Mapile, Maria Reynalen F. Institute of Biology, College of Science, University of the Philippines, Diliman, Quezon City 1101, Philippines Author Obusan, Maria Christine M. Institute of Biology, College of Science, University of the Philippines, Diliman, Quezon City 1101, Philippines Author Zhang, Yufeng 0000-0002-4066-6355 Hebei Key Laboratory of Animal Diversity, College of Life Science, Langfang Normal University, Langfang 065000, China Author Castañares, Edgar 0000-0002-3962-8837 College of Fisheries and Marine Sciences, Mindanao State University at Naawan, Naawan 9023, Misamis Oriental, Philippines Author Florida, Eric John D. College of Fisheries and Marine Sciences, Mindanao State University at Naawan, Naawan 9023, Misamis Oriental, Philippines & College of Environment and Life Sciences, Mindanao State University at Naawan, Naawan 9023, Misamis Oriental, Philippine Author Wu, Donghui Key Laboratory of Wetland Ecology and Environment, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, China & Department of Animal Science, Faculty of Tropical AgriSciences, Czech University of Life Sciences, Prague, Czech Republic text Zoosystematics and Evolution 2025 2025-02-21 101 1 389 404 journal article 308210 10.3897/zse.101.135702 21d82fde-8d6b-4301-bd49-0afdc7f49181 18C7202A-8137-44B2-A3D2-8C2DCCF58AD7

Table 3. Comparison among members of the Ph. dubia group of Sims and Easton (1972). The species are listed in order of size from largest to smallest.

Characters	Ph. callosa Gates, 1937	Ph. poiana Michaelsen, 1913	Ph. philippina Rosa, 1981	Ph. losbanosensis Aspe & Obusan, 2023	Ph. korinchiana Cognetti, 1922	Ph. vungtauensis Nguyen et al., 2018	Ph. dubia Horst, 1893	Ph. naawanensis sp. nov.	Ph. julkai Hong & James, 2011	Ph. banaoi Hong & James, 2010	Ph. lamaganensis Hong & James, 2011	Ph. balbalanensis Hong & James, 2010	Ph. globosa Hong & James, 2011
Length (mm)	330	290	180–240	220–228	83–180	132–169	80–140	82–100	49–69	53–60	46–55	42–50	35–42
Width (mm)	16	?	7	8–9	?	4.1 – 6.1	?	6 – 9	3.5–4	3–3.6	2.5–3.2	2.2–2.7	2.3–2.7
Pigmentation	?	gray	gray	brown	brown	grayinsh brown	brown	brown	light purple-brown	yellowish-brown	brown	reddish-brown	light brown
No. of segments	?	> 110	115–125	117–128	?	91 – 125	?	99 – 237	90–95	89–97	84–93	72–85	71–80
First dorsal pore	?	?	11 / 12	12 / 13	?	11 / 12 or 12 / 13	?	12 / 13	10 / 11	12 / 13	11 / 12	11 / 12	11 / 12
Setae on vii, xx	71, 114	50 (v); 52 (xix)	44,> 70	26–36, 55–60	38, 43	31–49 (viii), 57–82 (xxv)	38, 52	29–33, 69–71	29–31, 51–54	25–35, 46–47	43–44, 23–30	29–31, 41–42	29, 52
Setae bet. male pores	?	?	?	4–6	12	11–17	10	4–5	6–10	6–12	7–10	6–8	9
Dist. bet. sper. pores	?	0.4	5 th – 6 th setal lines	0.18	0.19–0.22	0.4	0.38	0.18	0.2–0.23	0.19	0.27–0.28	0.28–0.32	0.32
Dist. bet. male pores	?	0.29	7 th – 8 th setal lines	0.15	0.19–0.23	0.35	0.17	0.17	0.18–0.2	0.22	0.21–0.22	0.22	0.22
Caeca	?	?	xxvi – xxiii	xxvii – xxiii	?	xxvii – xxiv or xxiii	?	xxvii – xxiii	xxvii – xxv	xxvii – xxiv	xxvii – xxv	xxvii – xxiv	xxvii – xxvi
Prostate glands	?	?	xviii	xvii – xix	?	xvi – xviii	?	xvii – xix	xxvii – xxviii	xvi – xviii	xvii – xviii	xvii – xviii	xvii – xviii