Diversity and distribution of Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae) in Iran, with descriptions of three new species Author Hrivniak, Ľubos Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic & Faculty of Sciences, University of South Bohemia, Branisovska 31, 37005 Ceske Budejovice, Czech Republic lubos.hrivniak@gmail.com Author Sroka, Pavel Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic https://orcid.org/0000-0003-4367-6564 Author Bojkova, Jindriska Department of Botany and Zoology, Masaryk University, Kotlarska 2, 61137 Brno, Czech Republic Author Godunko, Roman J. Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic & Department of Invertebrate Zoology and Hydrobiology, University of Lodz, Banacha 12 / 16, 90237 Lodz, Poland https://orcid.org/0000-0003-2196-3327 Author Namin, Javid Imanpour Department of Fishery, Faculty of Natural Resources, University of Guilan, POB 1144, Sowmehsara-Rasht, Iran Author Bagheri, Samereh Department of Fishery, Faculty of Natural Resources, University of Guilan, POB 1144, Sowmehsara-Rasht, Iran Author Nejat, Farshad Department of Biodiversity and Ecosystem Management, Environmental Sciences Research Institute, Shahid Beheshti University, Daneshjou Boulevard, 1983969411 Tehran, Iran Author Abdoli, Ashgar Department of Biodiversity and Ecosystem Management, Environmental Sciences Research Institute, Shahid Beheshti University, Daneshjou Boulevard, 1983969411 Tehran, Iran Author Staniczek, Arnold H. Department of Entomology, State Museum of Natural History Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany https://orcid.org/0000-0001-5577-137X text ZooKeys 2020 947 71 102 http://dx.doi.org/10.3897/zookeys.947.51259 journal article http://dx.doi.org/10.3897/zookeys.947.51259 1313-2970-947-71 3297FBE4111C48499533225A53F7DB3C 8E93BC6985D85A52AD26A514847748FA

Table 1. List of records of the Caucasiron species found in Iran (three new species are not included). Abbreviations: RT - right tributary; LT - left tributary; JB - Jindriska Bojkova; TS - Tomas Soldan; IN - Javid Imanpour Namin; SB - Samereh Bagheri; AHS - Arnold H. Staniczek; MP - Milan Pallmann; RJG - Roman J. Godunko; FN - Farshad Nejat; AA - Ashgar Abdoli; HV - H. Valikhani; PT - P. Taban. Number of specimens includes larvae.

Species	Province	Stream	Locality	Nearest settlement	Altitude	Latitude (N) / Longitude (E)	Sampling date	Collector/reference	Number of specimens
E. (C.) iranicus	Alborz	Karaj R., Shahrestanak branch	SE of Shahrestanak	Chavar Chalun	2220	35°57'45.8"N, 051°21'59.7"E	1.9.2016	AA, HV, PT	4
	Ardabil	unnamed brook	in Alvaresi (below Alvares ski area)	Sarein	2235	38°09'38.0"N, 047°56'21.0"E	17.5.2016	JB, TS, IN; Bojkova et al. 2018	2
	Mazandaran	Koshk Sara R.	in Kosh Sara	Chalus	18	36°37'57.7"N, 051°28'04.4"E	25.9.2013	Mousavi and Hakobyan 2017	2
	Mazandaran	Firuz Abad R.	near Dasht Nazir	Marzan Abad	929	36°24'33.8"N, 051°24'42.5"E	11.9.2014	Mousavi and Hakobyan 2017	80
	Mazandaran	Haraz R.	in Gazanak	Gazanak	1590	35°54'08.3"N, 052°13'30.0"E	10.7.2013	Mousavi and Hakobyan 2017	8
	Mazandaran	Dalir R.	above Dalir	Marzan Abad	2126	36°19'23.2"N, 051°04'27.5"E	26.7.2014	Mousavi and Hakobyan 2017	174
	Mazandaran	Lasem R.	E of Polour	Polour	2180	35°50'04.1"N, 052°05'07.6"E	14.5.2017	AHS, MP, FN	1
	Mazandaran	RT of Sardab Rud	SW of Kelardasht	Kelardasht	2020	36°26'06.5"N, 051°03'52.6"E	8.3.2018	SB	85
	Mazandaran	LT of Sardab Rud	NW of Vandarbon	Kelardasht	2250	36°25'53.7"N, 051°01'59.1"E	9.3.2018	SB	63
	Mazandaran	Sardab Rud	S of Vandarbon	Kelardasht	2290	36°25'23.0"N, 051°02'12.4"E	10.3.2018	SB	5
	Tehran	Darban valley (type locality)	N of Tehran	Tehran	2100	35°50'24.0"N, 051°25'19.9"E	18.7.1970	Braasch and Soldan 1979	14
	Tehran	Lalan R.	above Zayegan	Fasham	2290	35°58'39.2"N, 051°34'56.5"E	8.5.2017	AHS, MP, RJG, FN	158
	Tehran	Lalan R.	in Lalan	Lalan	2440	35°59'50.3"N, 051°34'51.0"E	8.5.2017	AHS, MP, RJG, FN	87
	Tehran	Ahar R.	near Igol	Fasham	2020	35°55'11.2"N, 051°28'51.3"E	8.5.2017	AHS, MP, RJG, FN	1
	Tehran	Shahrestanak R.	NW of Shahrestanak	Asara	2100	35°59'01.2"N, 051°19'09.6"E	10.5.2017	AHS, MP, FN	1
E. (C.) nigripilosus	Alborz	Kordan R.	N of Kordan	Kordan	1430	35°57'15.6"N, 050°50'25.3"E	10.5.2017	AHS, MP, FN	4
	Mazandaran	RT of Dohezar R.	N of Holu Kaleh	Tonkaboon	880	36°37'37.5"N, 050°44'30.2"E	16.6.2018	SB	1
	Mazandaran	Dohezar R.	SW of Parde Sar	Tonkaboon	450	36°40'07.0"N, 050°49'20.0"E	16.6.2018	SB	1
	Tehran	Ahar R.	near Igol	Fasham	2020	35°55'11.2"N, 051°28'51.3"E	8.5.2017	AHS, MP, RJG, FN	5
	Tehran	Shahrestanak R.	NW of Shahrestanak	Asara	2100	35°59'01.2"N, 051°19'09.6"E	10.5.2017	AHS, MP, FN	1
E. (C.) cf. znojkoi	Alborz	Kordan R.	N of Kordan	Kordan	1430	35°57'15.6"N, 050°50'25.3"E	10.5.2017	AHS, MP, FN	4
	Gilan	RT of Khara Rud	S of Paein Khara Rud (S of Pashaki)	Sangar	210	37°02'29.0"N, 049°47'52.0"E	12.5.2016	JB, TS, IN	71
	Gilan	left fork of Khara Rud	in Madarsara (S of Pashaki)	Sangar	105	37°04'12.0"N, 049°46'36.0"E	12.5.2016	JB, TS, IN	27
	Gilan	right fork of Khara Rud	in Golestansara (S of Pashaki)	Sangar	201	37°02'20.0"N, 049°47'27.0"E	12.5.2016	JB, TS, IN	2
	Gilan	Zilaki River (RT of Sefid Rud)	in Mush Bijar (E of Shahr-e Bijar)	Shahr-e Bijar	120	37°00'28.0"N, 049°40'24.0"E	13.5.2016	JB, TS, IN	1
	Gilan	Sefidab (RT of Siah Rud)	in Divarsh (NE of Shirkuh)	Tutkabon	280	36°53'59.0"N, 049°35'06.0"E	13.5.2016	JB, TS, IN	149
	Gilan	Sangdeh (LT of Shafa Rud)	W of Punel	Punel	240	37°31'47.0"N, 049°00'52.0"E	15.5.2016	JB, TS, IN	10
	Gilan	Shafa Rud	W of Punel	Punel	240	37°31'47.0"N, 049°00'52.0"E	15.5.2016	JB, TS, IN	6
	Gilan	LT of Shafa Rud	NW of Sangdeh	Sangdeh	1345	37°31'46.0"N, 048°45'19.0"E	15.5.2016	JB, TS, IN	50
	Gilan	Shakhzar R.	NE of Fuman	Fuman	6	37°14'13.0"N, 049°20'43.0"E	15.5.2016	JB, TS, IN	1
	Gilan	LT of Bala Rud	S of Siahkal	Siahkal	490	37°00'31.0"N, 049°51'51.0"E	16.5.2016	JB, TS, IN	16
	Gilan	Lunak waterfalls	S of Siahkal	Siahkal	510	37°00'31.0"N, 049°51'49.0"E	16.5.2016	JB, TS, IN	5
	Gilan	Shamrud (RT of Sefid Rud)	S of Tushi (S of Siahkal)	Siahkal	315	37°03'00.0"N, 049°53'54.0"E	16.5.2016	JB, TS, IN	37
	Gilan	Chelavand R.	W of Chelvand	Lavandvil	-4	38°17'20.0"N, 048°51'35.0"E	19.5.2016	JB, TS, IN	2
	Gilan	unnamed brook	N of Chaldasht	Amlash	1255	36°59'33.0"N, 050°05'19.0"E	21.5.2016	JB, TS, IN	3
	Gilan	RT of Shalman Rud	in Bolurdekan	Amlash	345	37°01'09.0"N, 050°03'51.0"E	21.5.2016	JB, TS, IN	2
	Gilan	LT of Ghale Rudkhan	NE of Masuleh	Fuman	885	37°09'47.0"N, 049°00'17.0"E	22.5.2016	JB, TS, IN	66
	Gilan	RT of Ghale Rudkhan	NE of Masuleh	Fuman	705	37°09'42.0"N, 049°01'17.0"E	22.5.2016	JB, TS, IN	6
	Gilan	Ghale Rudkhan R.	E of Masuleh	Fuman	370	37°10'02.0"N, 049°05'03.0"E	22.5.2016	JB, TS, IN	4
	Golestan	unnamed river	in Shirinabad	Aliabad-e Katul	740	36°48'01.0"N, 055°01'05.0"E	27.4.2018	JB, TS, IN	50
	Golestan	Shirabad waterfalls	above Shirabad	Shirabad	140	36°57'33.0"N, 055°01'57.0"E	28.4.2018	JB, TS, IN	48
	Golestan	RT of Madarsu R.	E of Tangrah	Tangrah	495	37°23'27.0"N, 055°48'51.0"E	30.4.2018	JB, TS, IN	1
E. (C.) cf. znojkoi	Mazandaran	Shirinrud	S of Part Kola	Farim	770	36°09'02.5"N, 053°20'58.1"E	11.5.2017	AHS, MP, FN	203
	Mazandaran	trib. Kashpel R.	SW of Chamestan	Chamestan	400	36°25'31.4"N, 052°03'38.4"E	13.5.2017	AHS, MP, FN	17
	Mazandaran	Chelav R.	N of Pasha Kola	Pasha Kola	820	36°12'24.7"N, 052°25'51.9"E	14.5.2017	AHS, MP, FN	63
	Mazandaran	Chelav R.	NW of Pasha Kola	Pasha Kola	570	36°13'28.9"N, 052°23'35.4"E	14.5.2017	AHS, MP, FN	2
	Mazandaran	Baladeh R.	W of Razan	Razan	1360	36°11'39.6"N, 052°08'34.6"E	14.5.2017	AHS, MP, FN	9
	Mazandaran	Chai Bagh R.	E of Andar Koli	Ghaem Shahr	200	36°20'30.0"N, 052°54'03.0"E	8.5.2018	JB, TS, IN, SB	27
	Mazandaran	RT of Haraz R.	NW of Pasha Kola	Amol	570	36°13'27.0"N, 052°23'36.0"E	9.5.2018	JB, TS, IN, SB	5
	Mazandaran	LT of Haraz R.	in Panjab	Amol	955	36°05'52.0"N, 052°15'15.0"E	9.5.2018	JB, TS, IN, SB	1
	Mazandaran	unnamed brook	above Darab Kola	Neka	135	36°33'10.0"N, 053°15'32.0"E	10.5.2018	JB, TS, SB	2
	Mazandaran	unnamed brook	in Momey Khal	Ghaem Shahr	760	36°04'24.0"N, 052°58'19.0"E	11.5.2018	JB, TS, SB	1
	Mazandaran	Palang Darreh R.	SE of Shirgah	Shirgah	320	36°16'31.0"N, 052°56'54.0"E	11.5.2018	JB, TS, SB	2
	Mazandaran	LT of Palang Darreh R.	SE of Shirgah	Shirgah	345	36°16'30.0"N, 052°56'51.0"E	11.5.2018	JB, TS, SB	1
	Mazandaran	RT of Sardab Rud	SW of Kelardasht	Kelardasht	2020	36°26'06.5"N, 051°03'52.6"E	9.3.2018	SB	9
	Mazandaran	Sardab Rud	S of Vandarbon	Kelardasht	2290	36°25'23.0"N, 051°02'12.4"E	16.6.2018	SB	57
	Mazandaran	RT of Dohezar R.	N Holu Kaleh	Tonkabon	880	36°37'37.5"N, 050°44'30.2"E	16.6.2018	SB	83
	Mazandaran	Dohezar R.	SW Parde Sar	Tonkabon	450	36°40'07.0"N, 050°49'20.0"E	16.6.2018	SB	46
	Mazandaran	RT of Sehezar R.	S Parde Sar	Tonkabon	570	36°38'41.5"N, 050°50'11.1"E	16.6.2018	SB	20
	Mazandaran	Sehezar R.	S Parde Sar	Tonkabon	540	36°39'01.0"N, 050°50'00.0"E	24.8.2018	SB	21
	Mazandaran	Lavij Rud	SE of Kiakola	Noor	820	36°21'33.1"N, 052°03'11.0"E	24.8.2018	SB	8
	Mazandaran	Vaz Rud	E of Vaz Oliya	Noor	1140	36°19'08.0"N, 052°08'24.1"E	15.6.2018	SB	33
	Mazandaran	Safarud	SW of Ramsar	Ramsar	490	36°52'55.8"N, 050°33'56.1"E	15.6.2018	SB	1
	Mazandaran	LT of Safarud	SW of Ramsar	Ramsar	610	36°53'24.4"N, 050°33'56.1"E	15.6.2018	SB	2
	Mazandaran	LT of Safarud	SW of Ramsar	Ramsar	330	36°54'06.1"N, 050°35'12.1"E	15.6.2018	SB	11
	Mazandaran	Chalak Rud	SW of Galeshmahalleh	Ramsar	100	36°49'13.0"N, 050°43'23.8"E	15.6.2018	SB	3
	Mazandaran	LT of Chalak Rud	NW of Talesh Sara	Ramsar	180	36°50'46.9"N, 050°40'25.9"E	8.3.2018	SB	8
	Tehran	Ahar R.	near Igol	Fasham	2020	35°55'11.2"N, 051°28'51.3"E	8.5.2017	AHS, MP, RJG, FN	4