Taxonomic reassessment of salamanders (genus Hynobius) from Tsushima Islands Japan, with a resurrection of Hynobius tagoi Dunn, 1923 (Amphibia: Caudata) Author Niwa, Keita 0009-0009-6442-9555 Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Present affiliation: Akita Prefectural Office, Sanno 4 - 1 - 1, Akita 010 - 8570, JAPAN. & tsushimanda 48 @ gmail. com; https: // orcid. org / 0009 - 0009 - 6442 - 9555 tsushimanda48@gmail.com Author Nishikawa, Kanto 0000-0002-6274-4959 Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Present affiliation: Akita Prefectural Office, Sanno 4 - 1 - 1, Akita 010 - 8570, JAPAN. & Graduate School of Global Environmental Studies, Kyoto University, Yoshida Honmachi, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Faculty of Agriculture and Life Science, Hirosaki University, Bunkyo-cho 3, Hirosaki, Aomori 036 - 8561, JAPAN. kuroo @ hirosaki-u. ac. jp; https: // orcid. org / 0000 - 0003 - 0278 - 2758 & nishikawa. kanto. 8 v @ kyoto-u. ac. jp; https: // orcid. org / 0000 - 0002 - 6274 - 4959 nishikawa.kanto.8v@kyoto-u.ac.jp Author Matsui, Masafumi 0000-0003-2032-2528 Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Present affiliation: Akita Prefectural Office, Sanno 4 - 1 - 1, Akita 010 - 8570, JAPAN. & fumi @ zoo. zool. kyoto-u. ac. jp; https: // orcid. org / 0000 - 0003 - 2032 - 2528 fumi@zoo.zool.kyoto-u.ac.jp Author Kanamori, Sally 0000-0003-3798-2653 Graduate School of Global Environmental Studies, Kyoto University, Yoshida Honmachi, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Faculty of Agriculture and Life Science, Hirosaki University, Bunkyo-cho 3, Hirosaki, Aomori 036 - 8561, JAPAN. kuroo @ hirosaki-u. ac. jp; https: // orcid. org / 0000 - 0003 - 0278 - 2758 & s. kanamori. 5328 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3798 - 2653 s.kanamori.5328@gmail.com Author Kuro-O, Masaki 0000-0003-0278-2758 kuroo@hirosaki-u.ac.jp text Zootaxa 2023 2023-08-30 5339 3 201 236 http://dx.doi.org/10.11646/zootaxa.5339.3.1 journal article 265688 10.11646/zootaxa.5339.3.1 ff895335-e448-4de5-a14d-44be257c2306 1175-5326 8308966 6882297C-7C5B-428C-AABF-F671E0440674

TABLE 3. Measurements of H. tsuensis, Hynobius sp., and H. nebulosus in adult specimens. Mean±SD of SVL, medians of percentage ratios (R) of metric characters to SVL, ratios of some metric characters, CG, and LO. Ranges are shown in parentheses.

	H. tsuensis		Hynobius sp. (H. tagoi)		H. nebulosus
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	Male (n=25)	Female (n=13)	Male (n=28)	Female (n=13)	Male (n=53)	Female (n=10)
SVL	64.0±3.7	64.0±3.1	64.6±3.4	63.9±3.4	61.4±4.5	59.3±4.7
	(55.8–70.9)	(59.2–69.5)	(59.6–71.3)	(56.4–70.2)	(53.6–69.5)	(52.4–70.4)
RHL	24.4	23.3	24.0	23.5	24.5	22.2
	(23.1–26.7)	(22.2–24.4)	(22.8–26.5)	(22.5–26.1)	(22.3–26.8)	(20.6–24.9)
RHW	17.1	16.6	17.1	16.9	17.4	16.1
	(16.0–18.0)	(16.0–17.8)	(15.2–18.4)	(15.6–18.4)	(15.1–19.4)	(14.6–17.6)
RMXHW	17.8	16.9	17.8	17.4	18.1	16.8
	(16.5–19.6)	(16.0–18.5)	(15.9–19.1)	(15.8–18.6)	(15.8–20.6)	(15.2–17.9)
RLJL	11.5	11.4	11.6	11.4	12.3	11.1
	(10.6–13.2)	(10.1–12.0)	(10.6–13.1)	(10.0–12.4)	(10.1–14.3)	(9.6–11.9)
	Male (n=25)	Female (n=13)	Male (n=28)	Female (n=13)	Male (n=53)	Female (n=10)
RSL	6.9	6.6	6.7	6.4	6.4	6.0
	(5.8–7.5)	(6.2–7.0)	(6.1–7.4)	(5.9–6.8)	(5.2–7.2)	(5.4–6.6)
RIND	6.2	6.0	6.0	6.0	5.8	5.3
	(5.5–6.9)	(5.7–6.4)	(5.5–6.9)	(5.4–6.6)	(4.4–6.8)	(5.0–6.1)
RIOD	6.2	5.8	6.1	5.9	5.8	5.4
	(5.4–6.8)	(5.3–6.2)	(5.4–6.4)	(5.4–6.5)	(4.6–6.8)	(5.2–6.1)
RUEW	3.2	3.1	3.0	3.1	2.8	2.7
	(2.8–3.4)	(2.8–3.8)	(2.7–3.4)	(2.7–3.2)	(2.1–3.4)	(2.5–3.4)
RUEL	5.4	5.0	5.2	5.1	5.5	5.0
	(5.0–6.0)	(4.6–5.9)	(4.8–6.6)	(4.8–5.5)	(4.5–6.3)	(4.4–5.7)
RAGD	51.4	52.9	50.9	52.9	52.4	54.3
	(48.5–53.7)	(50.8–56.4)	(48.6–56.3)	(49.8–56.6)	(48.7–55.9)	(52.1–58.1)
RTRL	75.6	76.7	76.0	76.5	75.5	77.8
	(73.3–76.9)	(75.6–77.8)	(73.5–77.2)	(73.9–77.5)	(73.2–77.7)	(75.1–79.4)
RTAL	81.8 (n=23)	76.7 (n=11)	85.6 (n=24)	76.4	77.2 (n=22)	70.7
	(75.3–92.2)	(56.1–82.8)	(78.1–94.6)	(65.5–89.2)	(57.6–89.9)	(59.2–75.5)
RBTAW	10.9	9.7	10.6	9.8	11.5	10.7
	(9.0–12.2)	(8.4–11.3)	(8.5–12.5)	(8.7–10.8)	(9.5–13.2)	(9.1–12.4)
RMTAW	5.8	4.7	5.5	4.6	6.6 (n=51)	6.2
	(4.3–7.9)	(3.7–6.8)	(4.4–7.9)	(4.1–6.0)	(5.1–8.9)	(5.0–8.5)
RBTAH	8.1	7.4	8.3	7.5	9.8	8.9
	(7.0–10.9)	(6.6–8.8)	(6.9–10.1)	(6.7–8.1)	(8.0–11.5)	(7.5–11.0)
RMTAH	8.8	7.5	9.1	8.3	11.2 (n=51)	10.2
	(7.1–11.8)	(6.7–10.0)	(7.1–12.0)	(6.7–10.6)	(8.1–14.8)	(8.5–14.1)
RMXTAH	9.4	7.8	9.7 (n=27)	8.6	11.7 (n=51)	10.5
	(7.4–12.3)	(6.7–10.3)	(7.2–12.2)	(7.3–10.6)	(8.4–15.3)	(8.5–14.2)
RFLL	22.6	21.5	23.3	22.2	22.4	20.5
	(19.6–24.3)	(18.3–23.6)	(21.5–26.1)	(20.1–26.6)	(19.4–24.8)	(18.2–23.5)
RHLL	28.5	27.0	29.0	27.8	28.6	26.4
	(26.5–32.1)	(24.7–30.0)	(26.9–32.5)	(25.3–29.0)	(24.6–31.5)	(25.3–28.8)
R2FL	4.5	4.4	4.3	4.6	4.3	4.3
	(3.1–5.7)	(3.8–5.1)	(3.5–5.4)	(4.0–5.7)	(3.4–5.4)	(3.7–4.8)
R3FL	4.4	4.4	4.4	4.5	4.1	4.2
	(3.0–5.0)	(3.7–5.1)	(3.4–5.5)	(4.0–5.5)	(3.2–5.1)	(3.3–5.0)
R3TL	7.5	7.3	7.5	7.8	7.2	7.5
	(6.2–9.1)	(5.1–8.2)	(6.3–8.6)	(6.7–8.7)	(4.2–9.2)	(5.8–8.4)
R5TL	2.7	2.7	3.2	3.1	2.7	2.3
	(1.6–3.7)	(2.4–3.5)	(2.1–4.5)	(2.0–4.0)	(1.5–3.6)	(1.2–2.7)
RVTW	5.2	5.2	5.0	4.8	5.4	4.7 (n=9)
	(4.8–6.7)	(4.8–5.9)	(4.4–6.3)	(4.5–5.3)	(4.5–6.7)	(4.3–5.5)
	Male (n=25)	Female (n=13)	Male (n=28)	Female (n=13)	Male (n=53)	Female (n=10)
RVTL	5.5	5.4	5.1	5.0	5.5	4.8 (n=9)
	(5.1–6.7)	(5.1–5.9)	(4.5–6.9)	(4.6–5.6)	(4.3–7.4)	(4.4–5.7)
VTW/VTL	95.5	94.6	97.1	93.3	97.1	93.5 (n=9)
	(80.5–113.8)	(86.5–109.4)	(86.1–113.8)	(86.8–100.0)	(79.5–114.3)	(87.5–111.5)
5TL/3TL	36.7	34.8	41.5	42.0	36.4	32.9
	(22.2–49.0)	(32.6–54.5)	(30.9–65.0)	(28.3–47.8)	(21.4–55.0)	(15.6–39.0)
MTAW/	64.8	56.5	59.3	56.0	59.2 (n=51)	60.4
MTAH	(54.1–73.5)	(52.2–67.6)	(51.3–69.4)	(48.3–65.6)	(46.7–76.6)	(52.5–76.7)
CG	13	13	13	13	13	13
	(12–13)	(12–14)	(12–14)	(12–13)	(12–14)	(12–13)
LO	-2.0	-3.0	-1.5	-2.5	-2.0	-3.5
	(-3.5 to -1.0)	(-4.0 to -2.0)	(-3.0 to -0.5)	(-3.5 to -0.5)	(-3.5 to -1.0)	(-4.5 to -2.0)