Taxonomic reassessment of salamanders (genus Hynobius) from Tsushima Islands Japan, with a resurrection of Hynobius tagoi Dunn, 1923 (Amphibia: Caudata) Author Niwa, Keita 0009-0009-6442-9555 Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Present affiliation: Akita Prefectural Office, Sanno 4 - 1 - 1, Akita 010 - 8570, JAPAN. & tsushimanda 48 @ gmail. com; https: // orcid. org / 0009 - 0009 - 6442 - 9555 tsushimanda48@gmail.com Author Nishikawa, Kanto 0000-0002-6274-4959 Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Present affiliation: Akita Prefectural Office, Sanno 4 - 1 - 1, Akita 010 - 8570, JAPAN. & Graduate School of Global Environmental Studies, Kyoto University, Yoshida Honmachi, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Faculty of Agriculture and Life Science, Hirosaki University, Bunkyo-cho 3, Hirosaki, Aomori 036 - 8561, JAPAN. kuroo @ hirosaki-u. ac. jp; https: // orcid. org / 0000 - 0003 - 0278 - 2758 & nishikawa. kanto. 8 v @ kyoto-u. ac. jp; https: // orcid. org / 0000 - 0002 - 6274 - 4959 nishikawa.kanto.8v@kyoto-u.ac.jp Author Matsui, Masafumi 0000-0003-2032-2528 Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Present affiliation: Akita Prefectural Office, Sanno 4 - 1 - 1, Akita 010 - 8570, JAPAN. & fumi @ zoo. zool. kyoto-u. ac. jp; https: // orcid. org / 0000 - 0003 - 2032 - 2528 fumi@zoo.zool.kyoto-u.ac.jp Author Kanamori, Sally 0000-0003-3798-2653 Graduate School of Global Environmental Studies, Kyoto University, Yoshida Honmachi, Sakyo-ku, Kyoto 606 - 8501, JAPAN. Faculty of Agriculture and Life Science, Hirosaki University, Bunkyo-cho 3, Hirosaki, Aomori 036 - 8561, JAPAN. kuroo @ hirosaki-u. ac. jp; https: // orcid. org / 0000 - 0003 - 0278 - 2758 & s. kanamori. 5328 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3798 - 2653 s.kanamori.5328@gmail.com Author Kuro-O, Masaki 0000-0003-0278-2758 kuroo@hirosaki-u.ac.jp text Zootaxa 2023 2023-08-30 5339 3 201 236 http://dx.doi.org/10.11646/zootaxa.5339.3.1 journal article 265688 10.11646/zootaxa.5339.3.1 ff895335-e448-4de5-a14d-44be257c2306 1175-5326 8308966 6882297C-7C5B-428C-AABF-F671E0440674

TABLE 5. Frequencies of each body coloration of H. tsuensis and Hynobius sp., and the coloration of each holotype of H. bicolor and H. tagoi. Bold numbers show modal values.

	H. tsuensis		H. bicolor	Hynobius sp. (H. tagoi)		H. tagoi
......Continued on the next page
*: n = 27 P1: uniformly blackish or purplish brown without markings P2: blackish or purplish brown with yellowish markings or spots P3: yellow with blackish or purplish brown markings or spots P4: brown with dark stipples P5: dark brown with dark or blackish markings or spots
	Male (n=25)	Female (n=13)	Male (holotype)	Male (n=28)	Female (n=13)	Male (holotype)
C1: Yellow stripe(s) on the upper side of tail
present	21 (84.0%)	13 (100.0%)	1	7 (25.0%)	10 (76.9%)
indistinct	4 (16.0%)			15 (53.6%)	3 (23.1%)	1
absent				6 (21.4%)
C2: Yellow stripe(s) on the lower side of tail
present	15 (60.0%)	11 (84.6%)			3 (23.1%)
indistinct	8 (32.0%)	2 (15.4%)	1	15 (53.6%)	10 (76.9%)
absent	2 (8.0%)			13 (46.4%)		1
C3: Lateral color of the anterior half of the tail (L)
P1	6 (24.0%)	1 (7.7%)
P2	13 (52.0%)	9 (69.2%)
P3	3 (12.0%)	2 (15.4%)			3 (23.1%)
P4				27 (96.4%)	10 (76.9%)
P5	3 (12.0%)	1 (7.7%)		1 (3.6%)
C4: Lateral color of the anterior half of the tail (R)
P1	6 (24.0%)	1 (7.7%)
P2	13 (52.0%)	8 (61.5%)	1
P3	3 (12.0%)	3 (23.1%)			3 (23.1%)
P4				27 (96.4%)	10 (76.9%)	1
P5	3 (12.0%)	1 (7.7%)		1 (3.6%)
C5: Lateral color of the posterior half of the tail (L)
P1	24 (96.0%)	9 (69.2%)
P2	1 (4.0%)	4 (30.8%)
P3					3 (23.1%)
P4				26 (96.3%)*	10 (76.9%)
P5				1 (3.7%)*
C6: Lateral color of the posterior half of the tail (R)
P1	24 (96.0%)	9 (69.2%)	1
P2	1 (4.0%)	4 (30.8%)
P3					3 (23.1%)
P4				26 (96.3%)*	10 (76.9%)	1
P5				1 (3.7%)*
C7: Dorsal color
P1	2 (8.0%)
P2	11 (44.0%)	7 (53.8%)	1
P3	3 (12.0%)	5 (38.5%)			2 (15.4%)
	Male (n=25)	Female (n=13)	Male (holotype)	Male (n=28)	Female (n=13)	Male (holotype)
P4	1 (4.0%)			27 (96.4%)	11 (84.6%)	1
P5	8 (32.0%)	1 (7.7%)		1 (3.6%)
C8: Markings (spots) on the venter
present	10 (40.0%)	8 (61.5%)		18 (64.3%)	10 (76.9%)
absent	15 (60.0%)	5 (38.5%)	1	10 (35.7%)	3 (23.1%)	1