Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan Author Shimizu, So Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan & DC and Overseas Challenge Program for Young Researchers, Japan Society for the Promotion of Science, Tokyo, Japan & Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK https://orcid.org/0000-0002-5202-4552 parasitoidwasp.sou@gmail.com Author Broad, Gavin R. Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK https://orcid.org/0000-0001-7223-5333 Author Maeto, Kaoru Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan text ZooKeys 2020 990 1 144 http://dx.doi.org/10.3897/zookeys.990.55542 journal article http://dx.doi.org/10.3897/zookeys.990.55542 1313-2970-990-1 7B73642C278D40F89091B26213C9A704 9F51F78CD53F5005A305DE65494002C4

Table 2. Indices used in the present paper.

Indices		Formula
Head (Fig. 2B)
GOI	Geno-Orbital Index	maximum breadth of eye in profile [be] / maximum breadth of gena in same line [bg]
Fore wing (Fig. 4A, C)
AI	Alar Index	length of 1m-cu&M between 2m-cu and bulla [cd] / length of 2rs-m [ab]
CI	Cubital Index	length of CU between 1m-cu&M and 2cu-a [gf] / length of 2cu-a [fh]
DI	Discoidal Index	maximum vertical distance between CU (between 2cu-a and 2m-cu) and 1m-cu&M [k] / length of CU between 2cu-a and 2m-cu [fe]
ICI	Intercubital Index	length of 2rs-m [ab] / length of M between 2m-cu and 2rs-m [bc]
SDI	Second Discoidal Index	length of CU between 2cu-a and 2m-cu [ef] / length of CU between M&RS and 1m-cu&M [gi]
SI	Sinuousness Index	maximum length between 1m-cu&M and a straight line connecting the intersection of M, 2m-cu, and 1m-cu&M and the intersection of 1m-cu&M and CU [l] / distance between the intersection of M, 2m-cu, and 1m-cu&M [c] and the intersection of 1m-cu&M and CU [g] [j]
SRI	Second Recurrent Index	length of 2m-cu [ce] / length of CU between 2cu-a and 2m-cu [ef]
Hind wing (Fig. 4B)
NI	Nervellar Index	length of CU between M and cu-a [mn] / length of cu-a [no]
Metasoma (Fig. 5)
DMI	Dorsal Metasomal Index	length of dorsum of T2 [e] / length of dorsum of T3 [f]
PI	Petiolar Index	distance between base of T1 and anterior margin of spiracle [a] / distance between posterior margin of spiracle and apex of T1 [b]
THI	Thyridium Index	distance between anterior margin of T2 and anterior margin of thyridium [c] / maximum diameter of thyridium [d]