Barbodes klapanunggalensis, a new species of blind subterranean fish (Cypriniformes, Cyprinidae) from Klapanunggal karst area, West Java, Indonesia, with notes on its conservation Author Wibowo, Kunto 0000-0003-4465-3022 Museum Zoologicum Bogoriense, Research Center for Biosystematics and Evolution, National Research and Innovation Agency, Cibinong 16911, Jawa Barat, Indonesia Author Willyanto, M. Iqbal Indonesian Speleological Society, Bogor 16121, Jawa Barat, Indonesia Author Dharmayanthi, Anik Budhi 0000-0003-4635-6099 Museum Zoologicum Bogoriense, Research Center for Biosystematics and Evolution, National Research and Innovation Agency, Cibinong 16911, Jawa Barat, Indonesia Author Rahmadi, Cahyo 0000-0001-5928-9513 Museum Zoologicum Bogoriense, Research Center for Biosystematics and Evolution, National Research and Innovation Agency, Cibinong 16911, Jawa Barat, Indonesia Author Lumbantobing, Daniel Natanael 0000-0002-1458-1470 Species Obscura, Depok 16434, Jawa Barat, Indonesia text ZooKeys 2025 2025-02-24 1229 43 59 journal article 308245 10.3897/zookeys.1229.135950 41e5b741-f998-4e29-85a8-dfe4a0f27180 E301C9D7-7F48-4077-B17A-82F47904B67D

Table 1. Counts and measurements (expressed as percentages of standard length) of Barbodes klapanunggalensis sp. nov., B. binotatus, B. microps, and B. cf. microps.

	B. klapanunggalensis sp. nov.		B. binotatus	B. microps			B. cf. microps
	Holotype	Paratype	Non-types	Syntypes of Barbus microps			Non-types
	MZB. 26657	MZB. 26656	n = 21	BMNH 1845.6. 22.334	BMNH 1845.6. 22.335	BMNH 1845.6. 22.336	n = 14
Standard length (mm)	63.8	73.0	46.4–78.7	103.4	93.6	77.8	48.6–107.5
Counts
Dorsal-fin rays	iv, 8 ½	iii, 8 ½	iii – iv (iii), 8 ½ – 9 ½ (8 ½)	iii, 8 ½	iii, 8 ½	iii, 7 ½	iii, 8 ½
Anal-fin rays	iii, 5 ½	iii, 5 ½	iii, 5 ½	iii, 5 ½	iii, 5 ½	iii, 5 ½	iii, 5 ½
Pectoral-fin rays	15	14	14–16 (15)	15	15	15	13–16 (15)
Pelvic-fin rays	10	9	8–10 (9)	9	9	9	8–10 (9)
Principal caudal-fin rays	10 + 9	10 + 9	10 + 9	10 + 9	10 + 9	10 + 9	10 + 9
Pored lateral-line scales	24 + 1	22 + 1	22–26 (24) + 0–2 (2)	24 + 2	24 + 2	24 + 2	22–25 (23 or 24) + 0–2 (2)
Pre-dorsal scales	10	10	9–12 (10 or 11)	10	11	11	10 or 11
Transverse scales above lateral line	4 ½	4 ½	4 ½ – 5 ½ (4 ½)	4 ½	4 ½	4 ½	4 ½
Transverse (1 st) scales below lateral line	3 ½	3 ½	3–4 ½ (3 ½)	3 ½	3 ½	3 ½	3 ½
Transverse (2 nd) scales below lateral line	3 ½	3 ½	3 ½ – 4 ½ (3 ½)	3 ½	3 ½	3 ½	3 ½
Circumpeduncular scales	12	12	12	12	12	12	12
Gill rakers	ca. 12	ca. 12	10–13 (12)	13	11	11	11–13 (12)
Vertebrae	29	28	–	29	29	30	–
Measurements
Body depth	27.6	43.1	28.3–35.1 (31.0)	32.1	33.5	31.6	31.3–39.4 (36.3)
Body width	14.1	22.5	13.4–18.7 (15.9)	17.1	17.2	15.9	16.0–19.2 (17.6)
Head length	32.9	35.3	25.3–29.7 (27.3)	30.6	28.8	30.6	28.2–31.7 (30.2)
Dorsal head length	23.5	25.7	18.4–21.9 (20.1)	23.8	21.8	21.3	21.0–25.3 (22.4)
Head depth	21.4	27.6	19.8–23.3 (21.3)	22.3	21.1	21.3	21.8–24.1 (22.9)
Pre-dorsal fin length	53.0	56.6	51.5–56.5 (54.1)	52.6	57.0	54.8	54.8–61.7 (57.8)
Pre-pectoral fin length	32.6	33.6	24.5–27.5 (26.0)	30.2	26.7	29.3	26.0–29.7 (27.9)
Pre-anal fin length	73.6	77.2	69.7–74.7 (72.0)	72.6	72.1	72.3	72.0–79.3 (76.0)
Pre-pelvic fin length	54.0	59.6	46.7–51.0 (48.9)	49.7	48.9	49.9	48.6–54.9 (52.0)
Dorsal fin length	24.3	25.9	21.4–26.7 (23.9)	22.2	broken	broken	20.1–27.8 (24.2)
Dorsal-fin base length	15.1	16.4	14.1–17.2 (15.7)	15.1	14.9	15.9	14.3–17.4 (15.8)
Pectoral fin length	26.0	31.4	19.9–23.3 (21.5)	20.2	20.2	21.8	20.2–23.8 (22.5)
Pelvic fin length	21.5	24.4	17.2–20.5 (19.0)	19.0	18.2	18.9	18.3–21.4 (19.9)
Anal fin length	17.7	19.6	15.3–21.2 (17.6)	15.8	16.2	18.4	15.4–19.7 (17.4)
Anal-fin base length	9.7	11.8	9.1–11.2 (10.3)	8.7	10.0	9.1	8.9–12.1 (10.4)
Caudal peduncle depth	13.2	18.2	13.2–15.9 (14.7)	12.1	13.3	12.7	13.2–15.7 (14.4)
Caudal peduncle length	18.8	18.4	17.4–21.9 (20.0)	18.5	19.5	19.4	16.3–20.2 (18.0)