Solving the taxonomic identity of Hipposideros cineraceus sensu lato (Chiroptera: Hipposideridae) in the Thai-Malay Peninsula, with the description of a new species Author Wongwaiyut, Phutita 0009-0006-3416-3731 Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, Thailand, 90110. https: // orcid. org / 0009 - 0006 - 3416 - 3731 Author Karapan, Sunate Halabala Wildlife Research Station, Wildlife Research Division, Wildlife Conservation Bureau, Department of National Parks, Wildlife Author Saekong, Puchit Halabala Wildlife Research Station, Wildlife Research Division, Wildlife Conservation Bureau, Department of National Parks, Wildlife Author Francis, Charles M. 0000-0002-8648-3239 Canadian Wildlife Service, Environment and Climate Change Canada, Ottawa, Ontario, Canada, K 1 A 0 H 3. https: // orcid. org / 0000 - 0002 - 8648 - 3239 Author Guillén-Servent, Antonio 0000-0001-6273-485X Instituto de Ecología, A. C., Ctra. Antigua a Coatepec # 351, El Haya, 91073 Xalapa, Veracruz, México. https: // orcid. org / 0000 - 0001 - 6273 - 485 X Author Senawi, Juliana Department of Biological Sciences and Biotechnology, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, 43600 & Museum of Zoology, The Natural History Museum, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, 43600 & Langkawi Research Centre, Tuanku Abdul Halim Mu’adzam Shah Campus, Universiti Kebangsaan Malaysia, Jalan Teluk Yu, Teluk Author Khan, Faisal Ali Anwarali 0000-0001-7585-4885 Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, 94300 Kota Samarahan, Sarawak, Malaysia. https: // orcid. org / 0000 - 0001 - 7585 - 4885 Author Bates, Paul J. J. 0000-0003-3630-739X Harrison Institute, Bowerwood House, 15 St. Botolph’s Road, Sevenoaks, Kent, TN 13 3 AQ, United Kingdom. https: // orcid. org / 0000 - 0003 - 3630 - 739 X Author Jantarit, Sopark Excellence Center for Biodiversity of Peninsular Thailand, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, Author Soisook, Pipat text Zootaxa 2023 2023-05-03 5277 3 401 442 http://dx.doi.org/10.11646/zootaxa.5277.3.1 journal article 54346 10.11646/zootaxa.5277.3.1 7b4e6d3e-562e-4715-98db-a43eb92e2f60 1175-5326 7890018 83191D39-BF28-42ED-A31E-112FC3BB524C

TABLE 2. Cranial and dental measurements (in mm) of H. kingstonae sp. nov., H. einnaythu, H. cf. saevus, H. cf. cineraceus, H. kunzi, H. bicolor, H. ater, and H. cf. gentilis. See methods for the definition of measurements. Sample size, mean ± standard deviation (SD) are provided, with min–max in parentheses.

Characters	H. kingstonae sp. nov.		H. einnaythu		H. cf. saevus		H. cf. cineraceus		H. kunzi		H. bicolor		H. ater		H. cf. gentilis
	N	Mean±SD	N	Mean±SD	N	Mean±SD	N	Mean±SD	N	Mean±SD	N	Mean±SD	N	Mean±SD	N	Mean±SD
GTL	11	16.29±0.57	7	17.16±0.71	23	16.96±0.74	73	15.55±0.27	53	18.05±0.31	19	18.95±0.25	14	16.20±0.31	47	17.82±0.46
		(15.94–17.90)		(15.71–17.71)		(15.25–18.65)		(14.82–16.28)		(17.28–18.76)		(18.53–19.54)		(15.60–16.76)		(16.73–18.77)
SL	11	16.13±0.60	7	17.04±0.83	23	16.74±0.77	74	15.48±0.29		-	6	18.38±0.16	14	15.87±0.31	47	17.80±0.46
		(15.73–17.79)		(15.40–17.71)		(14.77–18.36)		(14.52–16.38)				(18.12–18.55)		(15.25–16.39)		(16.65–18.77)
CBL	11	14.37±0.67	7	15.13±0.60	22	14.98±0.90	73	13.63±0.38	53	15.66±0.27	19	16.62±0.26	13	14.20±0.33	47	15.88±0.45
		(13.95–16.30)		(13.90–15.64)		(12.65–16.98)		(12.86–15.83)		(15.20–16.33)		(16.26–17.20)		(13.47–14.73)		(15.01–16.72)
CCL	11	14.00±0.66	7	14.65±0.53	22	14.59±0.94	75	13.19±0.27	54	15.39±0.26	19	16.24±0.22	14	13.72±0.29	48	15.49±0.43
		(13.56–15.94)		(13.57–15.07)		(11.77–16.34)		(12.56–13.85)		(14.95–16.04)		(15.98–16.75)		(13.08–14.28)		(14.60–16.26)
ZB	11	8.01±0.16	7	8.30±0.41	20	8.23±0.25	77	7.40±0.22	54	9.10±0.17	19	9.19±0.25	13	8.08±0.16	48	8.95±0.24
		(7.79–8.26)		(7.49–8.64)		(7.83–8.7)		(6.79–7.95)		(8.63–9.60)		(8.64–9.73)		(7.86–8.37)		(8.33–9.38)
BB	11	7.49±0.17	7	7.92±0.52	23	7.65±0.38	77	7.07±0.19	54	8.23±0.20	19	8.48±0.25	14	7.05±0.16	48	8.33±0.29
		(7.19–7.78)		(6.96–8.31)		(6.89–7.89)		(6.54–7.59)		(7.70–8.61)		(8.05–9.08)		(6.84–7.54)		(7.55–8.78)
MW	11	8.32±0.12	7	8.61±0.20	22	8.51±0.17	78	7.79±0.17	54	9.05±0.12	19	9.29±0.14	13	8.32±0.14	48	8.95±0.18
		(8.07–8.50)		(8.32–8.75)		(8.18–8.87)		(7.45–8.26)		(8.79–9.30)		(9.12–9.62)		(8.10–8.58)		(8.62–9.35)
PC	11	2.62±0.14	7	2.53±0.19	23	2.76±0.22	78	2.57±0.12	54	2.68±0.12	19	2.95±0.10	13	2.48±0.12	48	2.70±0.13
		(2.40–2.91)		(2.28–2.87)		(2.52–3.44)		(2.33–2.86)		(2.47–3.05)		(2.81–3.17)		(2.30–2.74)		(2.38–2.89)
C-M3	11	5.41±0.18	7	5.69±0.32	23	5.66±0.28	75	5.05±0.15	55	6.14±0.12	19	6.41±0.10	15	5.29±0.20	49	6.11±0.20
		(5.12–5.81)		(5.05–5.99)		(4.71–6.17)		(4.65–5.43)		(5.90–6.46)		(6.19–6.71)		(4.72–5.60)		(5.71–6.46)
C1-C1	11	3.22±0.20	7	3.38±0.22	23	3.00±0.27	74	2.71±0.16	55	3.64±0.2	19	3.56±0.17	14	3.20±0.16	49	3.39±0.15
		(2.88–3.52)		(2.95–3.54)		(2.05–3.46)		(2.25–3.14)		(3.28–3.89)		(3.24–3.94)		(2.91–3.38)		(3.05–3.64)
M3-M3	11	5.54±0.15	7	5.77±0.31	7	5.70±0.33	78	5.01±0.19	55	6.16±0.17	19	6.14±0.13	15	5.48±0.19	49	6.10±0.23
		(5.32–5.83)		(5.20–6.06)		(5.19–6.41)		(4.59–5.56)		(5.38–6.49)		(5.84–6.37)		(5.23–5.85)		(5.66–6.69)
c-m3	11	5.67±0.32	7	6.02±0.37	23	5.96±0.31	77	5.28±0.23	54	6.58±0.13	19	6.83±0.11	15	5.73±0.22	49	6.50±0.25
		(5.05–6.28)		(5.26–6.33)		(5.07–6.61)		(4.43–5.88)		(6.23–6.91)		(6.65–7.10)		(5.36–6.09)		(5.92–7.11)
ML	11	9.64±0.45	7	10.34±0.60	19	10.40±0.54	75	8.95±0.27	54	11.28±0.21	19	11.76±0.20	13	9.76±0.24	49	10.93±0.33
		(9.22–10.92)		(9.17–10.83)		(8.98–11.49)		(8.06–9.70)		(10.72–11.82)		(11.46–12.17)		(9.45–10.16)		(10.18–11.76)
ALSW	11	4.29±0.13	7	4.37±0.16	23	4.27±0.23	79	3.96±0.12	54	4.47±0.10	19	4.79±0.10	14	4.11±0.08	48	4.54±0.10
		(4.04–4.43)		(4.12–4.61)		(3.80–4.59)		(3.64–4.31)		(4.13–4.67)		(4.56–4.98)		(3.96–4.28)		(4.23–4.85)
AMSW	10	3.07±1.04	7	2.47±0.12	19	2.61±0.27	79	2.40±0.29		-		-	12	2.66±0.24	48	2.41±0.22
		(1.99–4.28)		(2.16–3.25)		(2.29–3.20)		(2.01–3.80)						(2.37–3.08)		(2.05–2.85)