Four new species of polychromatic spiny-tailed iguanian lizards, genus Stenocercus (Iguania: Tropiduridae), from Peru Author Venegas, Pablo J. 0000-0002-6501-4492 Rainforest Partnership, 4005 Guadalupe St., Austin, Texas 78751, USA. & Instituto Peruano de Herpetología (IPH). Salazar Bondy 136, Santiago de Surco, Lima, Perú. & División de Herpetología-Centro de Ornitología y Biodiversidad (CORBIDI). Santa Rita No. 105 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, Perú. & sancarranca @ yahoo. es; https: // orcid. org / 0000 - 0002 - 6501 - 4492 sancarranca@yahoo.es Author García-Ayachi, Luis A. Instituto Peruano de Herpetología (IPH). Salazar Bondy 136, Santiago de Surco, Lima, Perú. & División de Herpetología-Centro de Ornitología y Biodiversidad (CORBIDI). Santa Rita No. 105 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, Perú. Author Chávez-Arribasplata, Juan C. Instituto Peruano de Herpetología (IPH). Salazar Bondy 136, Santiago de Surco, Lima, Perú. & División de Herpetología-Centro de Ornitología y Biodiversidad (CORBIDI). Santa Rita No. 105 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, Perú. Author García-Bravo, Antonio División de Herpetología-Centro de Ornitología y Biodiversidad (CORBIDI). Santa Rita No. 105 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, Perú. & Escuela de Ingeniería Forestal, Facultad de Ingeniería y Ciencias Agrarias, Universidad Nacional Toribio Rodríguez de Mendoza de Amazonas. Calle Higos Urco N 342 - 350 - 356. Amazonas, Perú. text Zootaxa 2022 2022-03-11 5115 1 1 28 journal article 20278 10.11646/zootaxa.5115.1.1 dc0b9aae-2fa1-4b3d-bc94-4cca9d764067 1175-5326 6345982 7B6B2504-7474-4624-93A1-5414D15C91BC

TABLE 1. Variation in scutellation and sexual dimorphism in snout–vent length (mm) of Stenocercus asenlignus sp. nov., S. nigrocaudatus sp. nov., S. leybachi sp. nov., and qalaywasi sp. nov., and their most similar species from the eastern slopes of the Central Andes, S. flagracanthus, S. torquatus, and S. crassicaudatus. Range followed by mean ± standard deviation is given for quantitative characters if applicable. The scale counts and measurements of S. torquatus, and S. crassicaudatus were taken from Torres-Carvajal (2007b) and for S. flagracanthus from Venegas et al. (2020a).

Character	S. asenlignus	S. nigrocaudatus	S. leybachi	S. qalaywasi	S. flagracanthus	S. torquatus	S. crassicaudatus
	n = 24	n = 9	n = 23	n = 2	n = 8	n = 46	n = 31
Scales around midbody	78–111 96.41±8.63	99–117 106.78±5.38	74–97 86.43±5.62	87–92 89.5	96–104 99.63±2.88	102–137 116.96±8.21	97–121 108.87±5.99
Vertebrals	71–106 89.08±9.15	96–108 101.78±3.77	63–73 68.65±3.20	77–79 78	83–97 89.38±5.21	83–115 98.86±7.94	83–97 89.80±3.74
Paravertebrals	92–118 105.87±7.72	110–128 120.89±6.17	100–112 105.22±3.32	104–108 106	95–111 100.13±5.41	103–151 124.05±12.17	107–166 126.67±12.21
Gulars	47–60 54.87±3.51	48–59 53.33±3.57	42–61 52.04±4.43	44–44 44	55–62 57.50±2.20	47–67 54.09±4.58	44–55 49.57±2.73
Supraoculars	5–8 6	5–7 6	7–9 8	6–6 6	4–6 5	6–8 7	6–8 6
Internasals	4–5 4	4–5 4	4–4 4	4–4 4	4–6 5	4–6 4	4–7 6
Subdigitals finger IV	23–30 27.45±1.76	26–31 27.33±1.58	21–28 24.00±1.48	26–29 27.25	26–29 27.38±1.19	22–29 25.76±1.49	23–32 28.53±1.80
Subdigitals toe IV	28–34 30.87±1.51	30–36 32.67±2.12	28–32 29.50±1.01	32–34 33	30–33 31.28±1.06	26–32 28.96±1.55	26–38 33.00±2.86
Tail length / Total lenght	0.49–0.57 0.53±0.02 n = 18	0.5–0.55 0.52±0.02 n = 7	0.49–0.58 0.55±0.03 n = 13	0.57–0.61 0.59 n = 2	0.51–0.54 0.52±0.01 n = 7	0.47–0.54 0.51±0.02 n = 26	0.57–0.62 0.59±0.01 n = 17
Maximum SVL males	73	72	84	88	77	84	95
Maximum SVL females	73	76	82	95	68	74	85
Black antehumeral collar in adult males	Present (incomplete midorsally)	Present (incomplete midorsally)	Absent	Present (complete midorsally)	Present (incomplete midorsally)	Present (complete midorsally)	Present (incomplete midorsally) or absent
Dorsal background color	Brown, gray, dark gray or green in the anterior half of dorsum	Dark gray, olive brown, or green	Dark gray, dark brown, or green	Gray, gray with bright green dots or green	Gray, grayish brown or pale brown	Dark brown, gray or emerald green	Dark brown or gray