A new species of the Boulenophrys (Anura, Megophryidae) from Hubei, China Author Li, Shize Department of Food Science and Engineering, Moutai Institute, Renhuai 564500, Guizhou, China & Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Chengdu, China Author Shi, Shengchao 0000-0003-2337-6572 Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Chengdu, China & Hubei Engineering Research Center for Protection and Utilization of Special Biological Resources in the Hanjiang River Basin, School of Life Science, Jianghan University, Wuhan 430056, Hubei, China Author Liu, Jing Department of Food Science and Engineering, Moutai Institute, Renhuai 564500, Guizhou, China Author Zhao, Jingjing Department of Food Science and Engineering, Moutai Institute, Renhuai 564500, Guizhou, China Author Gao, Shuo Nanjing Institute of Environmental Sciences, Ministry of Ecology and Environment of China, Nanjing 210042, Jiangsu, China Author Wang, Bin 0000-0001-6036-5579 Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Chengdu, China text Zoosystematics and Evolution 2025 2025-07-17 101 3 1213 1226 journal article 311134 10.3897/zse.101.155859 3d13c0a4-3203-40e0-8d9d-290077389356 37238A4D-4E12-4F86-A8E8-64362316C6A0

Table 2. References for morphological characters for congeners of the genus Boulenophrys.

ID	Species	References
1	B. acuta Wang, Li & Jin, 2014	Li et al. 2014
2	B. angka Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy & Che, 2019	Wu et al. 2019
3	B. anlongensis Li, Lu, Liu & Wang, 2020	Li et al. 2020
4	B. baishanzuensis Wu, Li, Liu, Wang & Wu, 2020	Wu et al. 2020
5	B. baolongensis Ye, Fei & Xie, 2007	Ye et al. 2007
6	B. binchuanensis Ye & Fei, 1995	Ye and Fei 1995; Lyu et al. 2023
7	B. binlingensis Jiang, Fei & Ye, 2009	Fei et al. 2009; Lyu et al. 2023
8	B. boettgeri Boulenger, 1899	Boulenger 1899; Lyu et al. 2023
9	B. brachykolos Inger & Romer, 1961	Inger and Romer 1961; Lyu et al. 2023
10	B. caobangensis Nguyen, Pham, Nguyen, Luong & Ziegler, 2020	Nguyen et al. 2020
11	B. caudoprocta Shen, 1994	Shen. 1994
12	B. cheni Wang & Liu, 2014	Wang et al. 2014
13	B. chishuiensis Xu, Li, Liu, Wei & Wang, 2020	Xu et al. 2020
14	B. congjiangensis Luo, Wang, Wang, Lu, Wang, Deng & Zhou, 2021	Luo et al. 2021
15	B. daiyunensis Lyu, Wang & Wang, 2021	Lyu et al. 2021
16	B. daoji Lyu, Zeng, Wang & Wang, 2021	Lyu et al. 2021
17	B. daweimontis Rao & Yang, 1997	Rao and Yang 1997
18	B. dongguanensis Wang & Wang, 2019	Wang et al. 2019 b
19	B. dupanglingensis Xiao & Mo, 2025	Xiao et al. 2025
20	B. elongata Zeng, Wang, Chen, Xiao, Zhan, Li & Lin, 2024	Zeng et al. 2024
21	B. fanjingmontis Zhang, Liang, Ran & Shen, 2012	Zhang et al. 2012
22	B. fansipanensis Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley, 2018	Tapley et al. 2018
23	B. fengshunensis Wang, Zeng, Lyu & Wang, 2022	Wang et al. 2022
24	B. frigida Tapley, Cutaja, Nguyen, Portway, Mahony, Nguyen, Harding, Luong & Rowley, 2021	Tapley et al. 2021
25	B. gaolanensis Song, Wang, Qi, Wang, Wang, 2024	Song et al. 2024
26	B. hengshanensis Qian, Hu, Mo, Gao, Zhang & Yang, 2023	Qian et al. 2023
27	B. hoanglienensis Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley, 2018	Tapley et al. 2018
28	B. hungtai Wang, Zeng, Lyu, Xiao & Wang, 2022	Wang et al. 2022
29	B. insularis Wang, Liu, Lyu, Zeng & Wang, 2017	Wang et al. 2017 a
30	B. jiangi Liu, Li, Wei, Xu, Cheng, Wang & Wu, 2020	Liu et al. 2020
31	B. jingdongensis Fei & Ye, 1983	Fei et al. 1983; Lyu et al. 2023
32	B. jinggangensis Wang, 2012	Wang et al. 2012
33	B. jiulianensis Wang, Zeng, Lyu & Wang, 2019	Wang et al. 2019 b
34	B. kuatunensis Pope, 1929	Pope 1929; Lyu et al. 2023
35	B. leishanensis Li, Xu, Liu, Jiang, Wei & Wang, 2018	Li et al. 2018
36	B. liboensis Zhang, Li, Xiao, Li, Pan, Wang, Zhang & Zhou, 2017	Zhang et al. 2017
37	B. lichun Lin, Chen, Li, Peng, Zeng, Wang, 2024	Lin et al. 2024
38	B. lini Wang & Yang, 2014	Wang et al. 2014
39	B. lishuiensis Wang, Liu & Jiang, 2017	Wang et al. 2017 b
40	B. lushuiensis Shi, Li, Zhu, Jiang, Jiang & Wang, 2021	Shi et al. 2021
41	B. minor Stejneger, 1926	Stejneger 1926; Lyu et al. 2023
42	B. mirabilis Lyu, Wang & Zhao, 2020	Lyu et al. 2020
43	B. mufumontana Wang, Lyu & Wang, 2019	Wang et al. 2019 b
44	B. nankunensis Wang, Zeng & Wang, 2019	Wang et al. 2019 b
45	B. nanlingensis Lyu, Wang, Liu & Wang, 2019	Wang et al. 2019 b
46	B. obesa Wang, Li & Zhao, 2014	Li et al. 2014
47	B. ombrophila Messenger & Dahn, 2019	Messenger et al. 2019
48	B. omeimontis Liu, 1950	Liu 1950; Lyu et al. 2023
49	B. palpebralespinosa Bourret, 1937	Bourret 1937; Lyu et al. 2023
50	B. pepe Wang & Zeng, 2024	Wang et al. 2024
51	B. puningensis Wang, Zeng, Lyu, Xiao & Wang, 2022	Wang et al. 2022
52	B. qianbeiensis Su, Shi, Wu, Li, Yao, Wang & Li, 2020	Su et al. 2020
53	B. rubrimera Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong & Rowley, 2017	Tapley et al. 2017
54	B. sangzhiensis Jiang, Ye & Fei, 2008	Jiang et al. 2008
55	B. sanmingensis Lyu & Wang, 2021	Lyu et al. 2021
56	B. shimentaina Lyu, Liu & Wang, 2020	Lyu et al. 2020
57	B. shuichengensis Tian & Sun, 1995	Tian and Sun 1995
58	B. shunhuangensis Wang, Deng, Liu, Wu & Liu, 2019	Wang et al. 2019 a
59	B. spinata Liu & Hu, 1973	Hu et al. 1973; Lyu et al. 2023
60	B. tongboensis Wang & Lyu, 2021	Lyu et al. 2021
61	B. tuberogranulatus Shen, Mo & Li, 2010	Mo et al. 2010
62	B. wugongensis Wang, Lyu & Wang, 2019	Wang et al. 2019 b
63	B. wuliangshanensis Ye & Fei, 1995	Ye and Fei 1995
64	B. wushanensis Ye & Fei, 1995	Ye and Fei 1995
65	B. xiangnanensis Lyu, Zeng & Wang, 2020	Lyu et al. 2020
66	B. xianjuensis Wang, Wu, Peng, Shi, Lu & Wu, 2020	Wang et al. 2020
67	B. xuefengmontis Lyu & Wang, 2023	Lyu et al. 2023
68	B. yangmingensis Lyu, Zeng & Wang, 2020	Lyu et al. 2020
69	B. yaoshanensis Qi, Mo, Lyu, Wang & Wang, 2021	Qi et al. 2021
70	B. yingdeensis Qi, Lyu, Wang & Wang, 2021	Qi et al. 2021
71	B. yunkaiensis Qi, Wang, Lyu & Wang, 2021	Qi et al. 2021