A taxonomic revision of the whitefish of lakes Brienz and Thun, Switzerland, with descriptions of four new species (Teleostei, Coregonidae) Author Selz, Oliver M. Department of Fish Ecology and Evolution, Centre for Ecology, Evolution & Biogeochemistry, Eawag: Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland & Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland https://orcid.org/0000-0002-2210-5909 oliver.selz@eawag.ch Author Doenz, Carmela J. Department of Fish Ecology and Evolution, Centre for Ecology, Evolution & Biogeochemistry, Eawag: Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland & Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland Author Vonlanthen, Pascal Department of Fish Ecology and Evolution, Centre for Ecology, Evolution & Biogeochemistry, Eawag: Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland & Aquabios GmbH, Les Fermes 57, 1792 Cordast, Switzerland Author Seehausen, Ole Department of Fish Ecology and Evolution, Centre for Ecology, Evolution & Biogeochemistry, Eawag: Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland & Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland text ZooKeys 2020 989 79 162 http://dx.doi.org/10.3897/zookeys.989.32822 journal article http://dx.doi.org/10.3897/zookeys.989.32822 1313-2970-989-79 F78F6D879DDB4CD98E4C60E4883A59B6 65AB71A5B3985F5E9FBF8CD56BC96143

Table 7. Morphological and meristic data of C. brienzii from Lake Brienz, Switzerland, NMBE-1077126, female, holotype; paratypes N = 12. For females and for both sexes the range and the mean include the holotype.

Morphological characters	C. brienzii	Lake Brienz
	Holotype	Both sexes
		N-total = 13		N-females = 4		N-males = 9
		Mean +/- Stdev	Range	Mean +/- Stdev	Range	Mean +/- Stdev	Range
SL (mm)	223.0	181.5 +/- 37.0	(118-226)	187.8 +/- 47.4	(118-223)	178.7 +/- 34.3	(129-226)
Percentage of standard length
PelvFB	4.1	3.7 +/- 0.6	(2.8-4.8)	3.6 +/- 0.6	(2.8-4.2)	3.8 +/- 0.6	(2.9-4.8)
PelvFS	6.1	6.1 +/- 0.8	(4.6-7.4)	5.8 +/- 0.4	(5.1-6.1)	6.3 +/- 0.9	(4.6-7.4)
PelvF	15.2	15.6 +/- 1.1	(14-17.5)	15 +/- 0.5	(14.6-15.7)	15.9 +/- 1.2	(14-17.5)
PecFB	3.1	3.1 +/- 0.2	(2.6-3.4)	3.0 +/- 0.3	(2.6-3.2)	3.1 +/- 0.2	(2.8-3.4)
PecF1	16.0	15.9 +/- 1.6	(13.9-20.1)	15.4 +/- 0.7	(14.5-16.0)	16.2 +/- 1.8	(13.9-20.1)
PecF2	17.0	16.8 +/- 1.6	(14-20.7)	16.4 +/- 0.7	(15.5-17)	17 +/- 1.9	(14-20.7)
DFB	12.3	11.8 +/- 0.8	(10.4-12.9)	11.7 +/- 0.4	(11.3-12.3)	11.9 +/- 1.0	(10.4-12.9)
DFAe	17.6	17.9 +/- 1.2	(15.5-19.8)	17.7 +/- 0.7	(16.9-18.6)	18.0 +/- 1.4	(15.5-19.8)
DFAd	18.7	18.6 +/- 1.5	(15.3-20.8)	18.3 +/- 0.4	(17.8-18.7)	18.8 +/- 1.8	(15.3-20.8)
DFPe	5.1	5.2 +/- 0.6	(4.2-6.5)	5.0 +/- 0.2	(4.7-5.2)	5.3 +/- 0.7	(4.2-6.5)
AFB	13.7	12.4 +/- 0.9	(11.1-13.7)	12.9 +/- 1.1	(11.4-13.7)	12.2 +/- 0.7	(11.1-13.6)
AFAe	11.1	11.2 +/- 1.0	(9.4-12.6)	11.2 +/- 0.5	(10.5-11.6)	11.2 +/- 1.2	(9.4-12.6)
AdFB	5.1	5.5 +/- 0.8	(4.0-7.1)	5.2 +/- 0.3	(5.0-5.7)	5.6 +/- 0.9	(4.0-7.1)
CF	23.8	24.1 +/- 1.1	(22.6-26.3)	23.2 +/- 0.5	(22.6-23.8)	24.5 +/- 1.1	(22.7-26.3)
CD	7.3	7.3 +/- 0.3	(6.7-7.7)	7.1 +/- 0.4	(6.7-7.5)	7.4 +/- 0.2	(7.1-7.7)
CL	13.9	13.8 +/- 1.0	(12.2-15.8)	14.0 +/- 0.4	(13.7-14.6)	13.7 +/- 1.1	(12.2-15.8)
PAdC	18.8	19.1 +/- 0.7	(17.9-20.7)	19 +/- 0.4	(18.6-19.4)	19.1 +/- 0.9	(17.9-20.7)
DHL	15.0	15.6 +/- 0.7	(14.6-16.8)	15.4 +/- 0.6	(15.0-16.3)	15.7 +/- 0.7	(14.6-16.8)
PreP	48.6	51.1 +/- 1.7	(47.8-54.0)	50.9 +/- 1.8	(48.6-52.8)	51.2 +/- 1.8	(47.8-54.0)
PreA	75.3	77.1 +/- 1.5	(74.3-79.5)	76.2 +/- 1.7	(74.3-78.2)	77.5 +/- 1.3	(75.4-79.5)
PreD	46.2	47.5 +/- 1.7	(43.9-49.4)	47.2 +/- 1.1	(46.2-48.2)	47.6 +/- 2	(43.9-49.4)
BD	24.6	22.6 +/- 1.7	(19.6-25.1)	22.7 +/- 2.5	(20.5-25.1)	22.6 +/- 1.5	(19.6-24.2)
PostD	45.9	44.1 +/- 1.1	(42.4-45.9)	44.6 +/- 1.3	(43-45.9)	43.9 +/- 1.1	(42.4-45.5)
TL	122.0	121.5 +/- 1.9	(117.8-124.4)	121.2 +/- 2.5	(117.8-123.8)	121.6 +/- 1.7	(119.2-124.4)
HL (mm)	45.4	38.7 +/- 7.3	(26.7-47.4)	39.2 +/- 8.5	(26.7-45.4)	38.5 +/- 7.3	(28.3-47.4)
Percentage of head length
SN	25.6	23.3 +/- 1.8	(20.5-26.3)	23.6 +/- 2.1	(21.1-25.6)	23.2 +/- 1.7	(20.5-26.3)
ED	24.4	25.3 +/- 1.6	(23.1-28.3)	25.2 +/- 1.6	(24.2-27.6)	25.3 +/- 1.7	(23.1-28.3)
EC	27.8	29 +/- 2.3	(25.6-32.9)	28.8 +/- 3.1	(25.6-32.9)	29.1 +/- 2.1	(26.5-32.7)
EH	22.0	24.4 +/- 1.4	(22-27.2)	23.9 +/- 1.7	(22-26.2)	24.7 +/- 1.3	(23-27.2)
ES	3.5	4.7 +/- 1.2	(3.3-7.2)	4.8 +/- 1.4	(3.5-6.5)	4.7 +/- 1.2	(3.3-7.2)
PostO	50.9	50.7 +/- 1.1	(48.2-52.3)	49.8 +/- 1.5	(48.2-51.1)	51.1 +/- 0.7	(50.3-52.3)
HD	75.2	68.5 +/- 3.3	(64.4-75.2)	69.8 +/- 4.4	(65.2-75.2)	67.9 +/- 2.8	(64.4-73.1)
MW	9.7	9.9 +/- 0.9	(8.5-10.9)	9.4 +/- 0.8	(8.5-10.3)	10.1 +/- 0.8	(8.6-10.9)
UJ	30.2	29.5 +/- 1.6	(27.1-32)	29 +/- 1.8	(27.1-30.8)	29.6 +/- 1.6	(27.3-32)
LJ	42.9	42.2 +/- 1.5	(40.5-45.7)	43.2 +/- 1.7	(42-45.7)	41.8 +/- 1.2	(40.5-43.7)
M	23.4	21 +/- 2.4	(15.4-24)	21 +/- 3.9	(15.4-24)	21.1 +/- 1.7	(18.3-23.8)
SD	7.0	8.8 +/- 1.4	(6.4-11.6)	7.4 +/- 0.9	(6.4-8.6)	9.4 +/- 1.2	(8-11.6)
SW	18.0	17.8 +/- 1.2	(15.7-20.2)	17.6 +/- 0.6	(16.7-18)	17.8 +/- 1.4	(15.7-20.2)
HW	52.1	48.1 +/- 3.1	(44.1-52.4)	48.5 +/- 4	(44.1-52.1)	47.9 +/- 2.9	(44.1-52.4)
IOW	28.4	26.2 +/- 1.9	(22.8-30.7)	26.3 +/- 1.7	(25-28.4)	26.1 +/- 2.1	(22.8-30.7)
INW	9.7	11.1 +/- 0.8	(9.7-12.6)	10.8 +/- 0.7	(9.7-11.2)	11.3 +/- 0.9	(10-12.6)
LJW	14.1	11.5 +/- 1.2	(10.1-14.1)	11.9 +/- 2	(10.1-14.1)	11.3 +/- 0.6	(10.5-12.4)
UJW	25.9	23.4 +/- 1.6	(20.2-26.1)	23.3 +/- 2.3	(20.2-25.9)	23.5 +/- 1.3	(21.4-26.1)
MGR	13.5	13.5 +/- 1.3	(10.9-15.1)	13.7 +/- 1.6	(11.6-15.1)	13.3 +/- 1.3	(10.9-14.9)
LGR	13.9	14.7 +/- 1.6	(12.1-16.8)	14.8 +/- 2.2	(12.1-16.8)	14.7 +/- 1.4	(13-16.7)
UA	20.4	18.5 +/- 1.7	(15.3-20.5)	19.6 +/- 0.8	(18.5-20.4)	18 +/- 1.7	(15.3-20.5)
LA	40.4	35.5 +/- 2	(33-40.4)	37.2 +/- 2.5	(35-40.4)	34.8 +/- 1.4	(33-37.5)
Meristic characters		Mode	Range	Mode	Range	Mode	Range
PelvF unbranched	1	1	(1-1)	1	(1-1)	1	(1-1)
PelvF branched	10	10	(9-11)	10	(9-10)	10	(10-11)
Meristic characters	C. brienzii	Lake Brienz
	Holotype	Both sexes
		N-total = 13		N-females = 4		N-males = 9
		Mode	Range	Mode	Range	Mode	Range
PecF unbranched	1	1	(1-1)	1	(1-1)	1	(1-1)
PecF branched	15	15	(15-17)	15	(15-17)	15	(15-17)
DF unbranched	4	4	(3-4)	4	(4-4)	4	(3-4)
DF branched	12	11	(10-13)	11	(10-12)	10	(10-13)
AF unbranched	4	4	(3-4)	4	(4-4)	4	(3-4)
AF branched	13	12	(11-13)	13	(11-13)	12	(12-12)
LS	89	86	(80-91)	89	(80-91)	86	(80-88)
PDS	36	35	(32-40)	na	(34-37)	32	(32-40)
TDS	9	9	(7-10)	9	(7-9)	9	(8-10)
TAS	8	8	(7-8)	8	(7-8)	8	(7-8)
TPS	8	8	(8-9)	8	(8-8)	8	(8-9)
UGR	14	14	(11-14)	13	(13-14)	12	(11-14)
LGR	25	24	(20-25)	24	(24-25)	23	(20-25)
total GR	39	37	(32-39)	37	(37-39)	32	(32-38)