Morphological and Genetic Evidence for the Stigmatodon goniorachis Complex (Tillandsioideae, Bromeliaceae) Reveals a New Species on Inselbergs in the Brazilian Atlantic Forest Author Manhães, Vitor da Cunha Programa de Pos-Graduaç ~ ao em Ci ^ encias Biologicas (Bot ^ anica), Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, S ~ ao Cristov ~ ao, 20940 - 040, Rio de Janeiro, RJ, Brazil & Instituto Nacional da Mata Atl ^ antica, Avenida Jose Ruschi, n 4, Centro, 29650 - 000, Santa Teresa, ES, Brazil & Authors for correspondence (manhaesvc @ gmail. com; afcosta @ acd. ufrj. br) manhaesvc@gmail.com Author Couto, Dayvid Rodrigues Instituto Nacional da Mata Atl ^ antica, Avenida Jose Ruschi, n 4, Centro, 29650 - 000, Santa Teresa, ES, Brazil Author Salgueiro, Fabiano Laboratorio de Biodiversidade e Evoluç ~ ao Molecular, Departamento de Bot ^ anica, Universidade Federal do Estado do Rio de Janeiro, Av. Pasteur 458, 22290 - 255, Rio de Janeiro, RJ, Brazil Author Costa, Andrea Ferreira da Departamento de Bot ^ anica, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, S ~ ao Cristov ~ ao, 20940 - 040, Rio de Janeiro, RJ, Brazil & Authors for correspondence (manhaesvc @ gmail. com; afcosta @ acd. ufrj. br) manhaesvc@gmail.com text Systematic Botany 2025 Basel, Switzerland 2025-01-06 49 4 673 694 https://doi.org/10.1600/036364424x17319705780145 journal article 10.1600/036364424X17319705780145 868f7e3f-9d8a-4760-8b46-874422c25ff9 1548-2324 15467112

TABLE 4. Morphological characters used to separate species of the Stigmatodon goniorachis complex.

Species
Morphological characters	S. carioca	S. goniorachis	S. costae	S. brassicoides	S. francae
Leaf blade length (cm)	16.8–30	19–30	18–24	10.7–16.5	30.5–41.3
Leaf blade width (cm)	1.7–3.7	1.4–3.0	2–4	1.5–2.8	2.3–3.8
Leaf blade trichomes	Densely lepidote,	Densely lepidote, not	Sub-densely lepidote,	Densely lepidote, not	Densely lepidote,
	forming white	forming white	not forming white	forming white	forming white
	crossbands	crossbands	crossbands	crossbands	crossbands
Longitudinal posture of	Revolute	Convolute	Convolute	Twisted to spiraled	Convolute
leaf blade
Inflorescence type	Simple	Simple, rarely	Simple	Simple	Compound, rarely
		compound			simple
Inflorescence length (cm)	12.4–24	12.7–38.2	20–30	12–17	81–125
Number of flowers	11 to 23	12 to 29	18 to 25	3 to 8	26–37
Peduncle position	Erect to suberect	Patent to pendulous	Erect, suberect to	Erect to suberect	Patent to pendulous
			recurved
Peduncle diameter (mm)	3–5	3.8–9	2.6–7	3.1–5	10–17
Peduncle bract dimensions	20–33 × 12–20	21–31 × 14–17	25–31 × 12–20	17–25 × 12–17	73–75 × 23–25
(mm)
Peduncle bracts 3	shorter than the	shorter or equal to the	shorter than the	shorter than the	exceeding the
internodes	internodes	internodes	internodes	internodes	internodes
Floral bract length (mm)	18–29	17–25	16–22	18–22	28–39
Sepal dimensions (mm)	20–28 × 13–18	22–29 × 13–17	20–23 × 12–19	21–33 × 14–16	28–35 × 18–26
Receptacle length (mm)	4–7	5–8	5–10	7–8	7.6–7.8(–18)