Tracing the footprints of the Atlas Brown Bear: a metric analysis of Holocene and Late Pleistocene remains from Northern Morocco Author Iken, Shaymae Instituto Universitario de Xeoloxía (IUX), Universidade da Coruña, Edificio de Servicios Centrales de Investigación (ESCI), Campus de Elviña s / n, 15071 A Coruña (Spain) & National Institute of Archaeological Sciences and Heritage (INSAP), Madinat Al Irfane, Hay Riad, BP 6828 Rabat Institutes (Morocco) Author García-Vázquez, Ana Instituto Universitario de Xeoloxía (IUX), Universidade da Coruña, Edificio de Servicios Centrales de Investigación (ESCI), Campus de Elviña s / n, 15071 A Coruña (Spain) & Archaeoscience Platform (AsP), Research Institute of the University of Bucharest (ICUB), University of Bucharest, 90 - 92 Sos. Panduri, 5 th District, 50663 Bucharest (Romania) Author González-Fortes, Gloria Instituto Universitario de Xeoloxía (IUX), Universidade da Coruña, Edificio de Servicios Centrales de Investigación (ESCI), Campus de Elviña s / n, 15071 A Coruña (Spain) Author Bouzouggar, Abdeljalil National Institute of Archaeological Sciences and Heritage (INSAP), Madinat Al Irfane, Hay Riad, BP 6828 Rabat Institutes (Morocco) & Max Planck Institute for Evolutionary Anthropology, Deutscher Platz 6, 04103 Leipzig (Germany) Author Grandal-D’Anglade, Aurora Instituto Universitario de Xeoloxía (IUX), Universidade da Coruña, Edificio de Servicios Centrales de Investigación (ESCI), Campus de Elviña s / n, 15071 A Coruña (Spain) aurora.grandal@udc.es text Comptes Rendus Palevol 2026 2026-03-11 25 4 57 76 https://doi.org/10.5852/cr-palevol2026v25a4 journal article 314543 10.5852/cr-palevol2026v25a4 0efbbe6b-2c87-4b27-9335-6ca8c35b0b01 1777-571X 19005282 urn:lsid:zoobank.org:pub:59E82E04-C7EB-4586-BF61-D9646DE136C6

TABLE 1. — Direct 14C dating of brown bear remains from several stratigraphic levels in Hattab II and of surface remains in Keft el-Hammar,and their 2σ calibration (95.4%) using software OxCal 4.4 (Bronk Ramsey 2009) and the IntCal20 calibration curve from Reimer et al. (2020).

Cave	Level	Bone code	Bone type	Dating lab code	AMS 14 C y BP	Cal y BP from	Cal y BP to
Keft el-Hammar	surface	HST-1	humerus	Beta-614566	9080 ± 30	10281	10189
Hattab II	Level 3	HAT-212	radius	Beta-614563	9400 ± 30	10716	10512
Hattab II	Level 8	HAT2-674	3rd phalanx	Beta-614564	9480 ± 30	11065	10584
Hattab II	Level 9	HAT2-1213	radius	Beta-614565	8850 ± 30	10155	9763