Two new cave-dwelling species of Triplophysa (Cypriniformes, Nemacheilidae) from Guizhou, China
Author
Zhu, Ting-Ting
0000-0003-4178-2785
School of Life Sciences, Guizhou Normal University, Guiyang 550025, China
Author
Yuan, Feng-Hua
https://orcid.org/0009-0004-5112-0683
School of Life Sciences, Guizhou Normal University, Guiyang 550025, China
Author
Zhang, Ren-Yi
0000-0002-3647-1416
School of Life Sciences, Guizhou Normal University, Guiyang 550025, China
Author
Zhao, Ya-Hui
0000-0002-4615-596X
State Key Laboratory of Animal Biodiversity Conservation and Integrated Pest Management, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China
text
ZooKeys
2026
2026-06-03
1281
123
145
journal article
315338
10.3897/zookeys.1281.186268
dbe69cd0-d13d-45cc-9840-03a3c7b03a16
AF3BE807-A4C1-4C5D-B590-4615C27C5124
Table 3. Morphology of two new species described in this paper was compared with that of 44 known species of hypogean group of Triplophysa ID Species Skin pigmentation Eye diameter (% HL ) Vertebrae Posterior chamber of air bladder Dorsal fin rays Anal fin rays Caudal fin rays CPL / CPD Ventral fin tip reaching to anus 1 T. aluensis Absent 5.6 – Degenerated iii, 7 iii, 5 13 2.8 No 2 T. anlongensis Whole body 5.1–9.3 4 + 37 Degenerated iii, 8 iii, 5 16 1.7 No 3 T. anshuiensis Dorsal Absent – Developed iv, 7–8 ii, 6 14 1.59 Yes 4 T. baishuijiangensis Absent 2.9–6.1 4 + 34 Developed iii, 7 ii, 5 15–16 2.3–2.7 Yes 5 T. baotianensis Whole body 14.0–15.0 – Degenerated iii, 6–7 ii, 5 11–13 1.7–2.3 No 6 T. cehengensis Absent 1.5–2.2 4 + 35 Developed iv, 9 iii, 5 16 2 Yes 7 T. erythraea Absent Absent – Developed ii, 8 i, 6 17 1.9–2.4 Yes 8 T. fengshanensis Absent Absent – – ii, 8 ii, 6 16 2.4–2.6 No 9 T. flavicorpus Whole body 5.1–6.8 4 + 34 Degenerated iii, 10 iii, 6–7 16 1.7 Yes 10 T. gejiuensis Absent Absent – Developed iii, 7–8 iii, 4–6 14–15 1.4 Yes 11 T. guizhouensis Whole body 9.4–12.1 – Developed iii, 8 iii, 6 14 1.7 No 12 T. huapingensis Whole body 10.4–14.3 – Degenerated iii, 8–9 iii, 5 16 1.7–1.8 No 13 T. langpingensis Absent 2.7–5.9 – – iii, 7–8 iii, 5–6 14 1.55 Yes 14 T. longipectoralis Whole body 11.8–16.4 4 + 35 Degenerated iii, 8 iii, 5–6 14–15 1.4 Yes 15 T. longliensis Whole body 9.5–11.5 4 + 38 Developed iii, 8 iii, 5 15–16 2.43 Yes 16 T. luochengensis Whole body 7.5–8.6 4 + 33–34 Degenerated iii, 8 ii, 6 16–17 1.6 No 17 T. macrocephala Whole body 3.6–8.0 – Degenerated iii, 7–9 iii, 5–6 15–17 1.95 Yes 18 T. nandanensis Whole body 11.1–21.3 4 + 36 Degenerated iv, 8 iv, 5 14–16 1.69 No 19 T. nanpanjiangensis Whole body 12.0–16.5 4 + 38 Degenerated iii, 7–8 ii, 5 16 1.7–2.3 No 20 T. nasobarbatula Whole body 9.1–13.3 4 + 36 Degenerated iii, 8 iii, 5 15 1. 4–1.8 Yes 21 T. nayongensis Absent Absent 4 + 35 Developed iii, 8 iii, 5–6 12–14 2.3–2.9 Yes 22 T. panzhouensis Whole body 7.0–11.0 4 + 35 Degenerated iv, 7–8 iii, 5 16 2.2 No 23 T. posterodorsalus Absent Absent – – iii, 6 ii, 4 15 3.33 No 24 T. qingzhenensis Whole body 2.1–4.4 4 + 36 Degenerated iii, 7–8 iii, 5 14 2.25 No 25 T. qini Absent Absent 4 + 34–35 – ii, 8 ii, 5 14–16 2.36 Yes 26 T. qiubeiensis Absent Absent 4 + 35 Degenerated iii, 7 iii, 5 14–15 2.0–2.8 Yes 27 T. rongduensis Whole body 7.2–14.7 4 + 39 Degenerated iv, 9 iii, 5 16 1.62 No 28 T. rosa Absent Absent – – iii, 9 iii, 6 14 2.3 Yes 29 T. sanduensis Whole body 11.9–15.4 4 + 37 Degenerated ii, 8–9 i, 5 17–18 1.58 No 30 T. shilinensis Absent Absent – Degenerated iii, 7 iii, 5 14 2.47 No 31 T. tianeensis Whole body 3.0–5.9 4 + 35 Degenerated iii, 6–7 iii, 6 15–16 1.84 No 32 T. tianlinensis Absent Absent – Degenerated iv, 8–9 iii, 6 15–16 1.55 Yes 33 T. tianxingensis Whole body 4.2–6.7 4 + 38 Developed iii, 8 ii, 5 16 2 No 34 T. wenshanensis Whole body 5.1–6.4 4 + 38 ~ 39 Normally developed iii, 8–10 iii, 6–8 17–19 2.2–2.8 No 35 T. wudangensis Whole body 5.1–6.5 4 + 34 Degenerated iii, 7 iii, 5 14 2.56 No 36 T. xiangshuingensis Whole body 7.5 – Degenerated iii, 6 iii, 5 14 1.9 No 37 T. xiangxiensis Absent Absent – Developed iii, 8 iii, 6 16 2.37 Yes 38 T. xichouensis Whole body Absent 4 + 36 Developed iii, 8 ii, 6 16 2.8 Yes 39 T. xingyiensis Whole body 9.1–17.3 4 + 36 Degenerated ii, 7 i, 5 16 1.9–2.2 No 40 T. xiuwenensis Absent 4.6–6.7 4 + 37 Degenerated iii, 8 iii, 6 14–15 2.1–2.5 No 41 T. xuanweiensis Absent Absent – Well developed iii, 7–8 iii, 5 17–18 2.02 Yes 42 T. yaluwang Whole body 4.6–6.1 4 + 36 Degenerated iii, 7 iii, 5 14 2.5 Yes 43 T. yangi Absent 2–5.3 4 + 34 ~ 35 Well developed iii, 7–9 iii, 5–7 15–18 1.7–1.9 Yes 44 T. yunnanensis Whole body 7.2–8.3 – Degenerated iii, 7 iii, 5 15–16 1.8–2.1 No 45 T. zhenfengensis Whole body 7.1–16.7 4 + 36 Degenerated iii, 7 iii, 5 14–15 1.6–2.0 No 46 T. ziyunensis Whole body 2.4–4.9 4 + 35 Degenerated iii, 8 iii, 5 16 2.34 Yes 47 T. dafangensis sp. nov. Absent 5.3–7.1 4 + 39 Degenerated iii, 7–8 iii, 6–7 14–16 2.3–2.8 Yes 48 T. zhijinensis sp. nov. Present Absent or 1.3–2.9 4 + 35 Degenerated iii, 8 iii, 5–6 14–15 2.0–2.9 No