High alpine sorcerers: revision of the cave wētā genus Pharmacus Pictet & de Saussure (Orthoptera: Rhaphidophoridae: Macropathinae), with the description of six new species and three new subspecies Author Hegg, Danilo 34DFC18A-F53D-417F-85FC-EF514F6D2EFD 135 Blacks Road, Opoho, Dunedin 9010, New Zealand. danilo_hegg@hotmail.com Author Morgan-Richards, Mary 48F2FB1A-4C03-477C-8564-5417F9739AE1 Wildlife & Ecology Group, Massey University, Private Bag 11 - 222, Palmerston North 4442, New Zealand. M.Morgan-Richards@massey.ac.nz Author Trewick, Steven A. 7A378EE1-BADB-459D-9BAA-7059A675F683 Wildlife & Ecology Group, Massey University, Private Bag 11 - 222, Palmerston North 4442, New Zealand. s.trewick@massey.ac.nz text European Journal of Taxonomy 2022 2022-04-04 808 1 1 58 http://dx.doi.org/10.5852/ejt.2022.808.1721 journal article 54975 10.5852/ejt.2022.808.1721 3eba97d9-c3b4-4437-8922-d01358c276bb 2118-9773 6419368 7898E29D-1F57-4DC2-AB70-8532234CA118

Table 1 (continued on next three pages). Dimensions and spine count of species of Pharmacus Pictet & de Saussure, 1893.

		Pharmacus montanus	Pharmacus cochleatus	Pharmacus cochleatus
			cochleatu s	nauclerus subsp. nov.
Sample size		27 (12 ♀♀, 15 ♂♂)	56 (24 ♀♀, 32 ♂♂)	10 (6 ♀♀, 4 ♂♂)
Apical spines fore, mid and hind femora 1		0 0, 0 1, 0 0	0 0, 0 1, 0 0	0 0, 0 1, 0 0
Apical spines fore, mid and hind tibiae		4-4-8 2 ♂♂: 4-4-9	4-4-8	4-4-8
Body length (mm) 2		♀ 15.0 (13.1–18.5) ♂ 14.4 (13.1–20.0)	♀ 14.4 (12.1–17.4) ♂ 12.6 (11.1–14.9)	♀ 15.3 (14.1–15.5) ♂ 12.3 (11.9–13.2)
Pronotum length (mm)		♀ 4.3 (3.8–4.9) ♂ 4.2 (3.5–4.6)	♀ 3.8 (2.8–4.8) ♂ 3.4 (2.5–4.2)	♀ 3.6 (3.3–4.2) ♂ 3.3 (2.5–3.5)
Eye colour		brown / black	brown / black	brown / black
Ovipositor length (mm)		10.2 (9.4–11.3)	10.3 (8.0–12.1)	10.1 (9.5–11.3)
Ratio ovipositor to body length		0.70 (0.52–0.78)	0.69 (0.60–0.87)	0.69 (0.63–0.75)
Teeth: ventral valve of ovipositor		7 (5–8)	7 (5–9)	7.5 (7–9)
Teeth: dorsal valve of ovipositor		0	0	0
Length of hind tibia (mm)		♀ 15.5 (12.3–18.1) ♂ 17.8 (14.6–18.5)	♀ 12.6 (10.4–16.3) ♂ 12.0 (10.0–15.4)	♀ 12.8 (11.1–14.5) ♂ 11.7 (11.0–11.9)
Ratio hind tibia to body length		♀ 1.04 (0.83–1.30) ♂ 1.18 (0.90–1.35)	♀ 0.86 (0.77–1.09) ♂ 0.94 (0.78–1.17)	♀ 0.88 (0.73–0.94) ♂ 0.93 (0.90–0.97)
Superior spines hind tibia	prolateral retrolateral	23 (17–32) 22 (16–28)	23 (14–29) 20 (14–29)	19.5 (18–22) 18 (16–21)
Spine density on hind tibia (count/ mm)	prolateral retrolateral	♀ 1.42 (1.22–1.62) ♂ 1.36 (1.09–1.78) ♀ 1.44 (1.15–1.55) ♂ 1.24 (0.96–1.45)	♀ 1.75 (1.16–2.31) ♂ 1.95 (1.09–2.55) ♀ 1.54 (1.13–2.31) ♂ 1.73 (1.09–2.36)	♀ 1.50 (1.31–1.89) ♂ 1.79 (1.51–1.91) ♀ 1.31 (1.24–1.71) ♂ 1.64 (1.43–1.76)
Pairs of longer spines on hind tibia		0	0	0
Superior spines on 1st tarsus segment		0	5 (1–9)	5 (4–7)
Superior spines on 2nd tarsus segment		0	0 (0–2)	0
Fore tibia, inferior spines	prolateral retrolateral	2 (1–3) 2 (1–2)	2 (1–2) 2 (1–2)	2 (2–2) 2 (2–2)
Fore tibia, superior spines		0	0	0
Mid tibia, inferior spines	prolateral retrolateral	2 (2–3) 2 (1–2)	2 (2–3) 2 (1–3)	2 (2–2) 2 (2–2)
Mid tibia, superior spines	prolateral retrolateral	0	0 0 (0–2)	0 0 (0–1)
Fore femur, Inferior spines	prolateral retrolateral	0	0	0
Mid femur, inferior spines	prolateral retrolateral	0	0	0
Hind femur, inferior spines	prolateral retrolateral	0 (0–1) 1 (0–2)	2 (0–3) 4 (0–5)	2 (1–2) 3.5 (2–4)