Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data Author Luo, Tao 0000-0003-4186-1192 School of Life Sciences, Yunnan University, Kunming, China & School of Karst Science, Guizhou Normal University, Guiyang, China & Southwest United Graduate School, Kunming, China Author Mao, Ming-Le 0000-0001-9764-4731 School of Karst Science, Guizhou Normal University, Guiyang, China Author Lan, Chang-Ting https://orcid.org/0009-0007-2381-3601 School of Karst Science, Guizhou Normal University, Guiyang, China Author Zhao, Zi-Fa https://orcid.org/0009-0000-1301-3159 School of Karst Science, Guizhou Normal University, Guiyang, China Author Wang, Zhong-Lian School of Karst Science, Guizhou Normal University, Guiyang, China Author Yu, Jing https://orcid.org/0009-0004-3629-3826 School of Karst Science, Guizhou Normal University, Guiyang, China Author Wang, Jia-Jia School of Karst Science, Guizhou Normal University, Guiyang, China Author Yan, Chen-Rui School of Karst Science, Guizhou Normal University, Guiyang, China Author Xiao, Ning 0000-0002-7240-6726 School of Life Sciences, Guizhou Normal University, Guiyang, China Author Zhou, Jiang 0000-0003-1560-8759 School of Karst Science, Guizhou Normal University, Guiyang, China text Zoosystematics and Evolution 2025 2025-06-02 101 3 1023 1055 journal article 310152 10.3897/zse.101.144375 7071d835-32b1-4289-a3b7-0c2beebe2610 798F3D1F-E08C-49AA-8662-30CBC75BF53F

Table 4. External and craniodental measurements (in mm) of M. milinensis sp. nov., M. yadongensis sp. nov., and four closely related species. MC: M. chrysochaetes (Eger et al. 2011); MH: M. harpioloides (Kruskop and Eger 2008); MY: M. yushuensis (Wang et al. 2024 a). NA indicates that the data is not available.

Species	M. yadongensis sp. nov.		M. milinensis sp. nov.		MC	MH	MY	M. yuanyang (Mou et al. 2024)
	Male (n = 3)	Female (n = 1)	Male (n = 6)	Female (n = 4)	Holotype (male)	Holotype (male)	Holotype (male)	Holotype (male)	Male (n = 2)	Female (n = 5)
BW	5.68–7.00 (6.52 ± 0.73)	6.86	3.15–3.49 (3.37 ± 0.12)	3.35–7.50 (4.73 ± 1.88)	NA	NA	NA	4.20	3.4–3.6	3.8–4.7 (4.3 ± 0.3)
HB	28.68–35.54 (32.21 ± 3.43)	33.88	33.70–37.89 (35.78 ± 1.57)	29.23–39.29 (34.30 ± 5.60)	NA	35.00	30.44	35.05	31.37–35.94	32.11–36.34 (34.82 ± 1.52)
TL	28.32–30.46 (29.52 ± 1.09)	32.12	25.54–30.32 (28.88 ± 1.69)	24.51–28.98 (26.66 ± 2.29)	28.00	30.50	28.08	31.58	23.89–27.25	26.92–31.58 (28.39 ± 1.64)
EL	11.40–14.02 (12.85 ± 1.33)	14.2	12.48–14.58 (13.46 ± 0.82)	13.12–13.95 (13.55 ± 0.34)	11.00	12.30	11.08	12.99	12.77–13.4	12.27–14.77 (13.25 ± 0.82)
EW	6.20–7.06 (6.54 ± 0.46)	8.04	6.92–8.47 (7.70 ± 0.52)	6.76–8.58 (7.38 ± 0.84)	NA	NA	NA	NA	NA	NA
TRL	7.15–7.49 (7.37 ± 0.19)	7.81	6.59–7.82 (7.31 ± 0.51)	6.28–8.35 (7.22 ± 1.04)	NA	NA	NA	NA	NA	NA
TRW	2.14–2.15 (2.14 ± 0.01)	2.13	1.51–5.35 (2.49 ± 1.42)	1.76–2.34 (2.03 ± 0.28)	NA	NA	NA	NA	NA	NA
HFL	6.07–8.55 (7.15 ± 1.27)	7.06	5.31–8.89 (7.37 ± 1.32)	6.82–9.18 (7.94 ± 0.97)	7.00	NA	8.46	7.88	6.14–6.83	6.25–8.52 (7.66 ± 0.75)
FL	28.81–30.29 (29.60 ± 0.75)	31.84	28.84–30.43 (29.82 ± 0.60)	30.83–34.01 (32.12 ± 1.35)	26.35	29.70	31.34	29.80	NA	NA
TIB	13.57–14.27 (13.82 ± 0.39)	14.61	12.64–13.95 (13.42 ± 0.60)	13.32–14.37 (13.94 ± 0.44)	10.92	NA	15.22	13.48	12.56–13.16	12.37–13.57 (13.15 ± 0.43)
GTL	13.73–14.11 (13.92 ± 0.27)	NA	14.07–14.27 (14.17 ± 0.14)	14.16	14.05	NA	14.14	14.13	13.44–14	13.95–14.16 (14.09 ± 0.07)
CCL	12.61–12.89 (12.75 ± 0.20)	NA	12.01–13.29 (12.65 ± 0.91)	12.26	12.45	12.34	13.85	12.12	11.58–11.93	11.94–12.22 (12.10 ± 0.09)
BCW	6.36–6.70 (6.53 ± 0.24)	NA	6.81–7.03 (6.92 ± 0.16)	6.92	6.98	7.21	7.23	6.95	6.91–6.93	6.93–7.15 (7.00 ± 0.08)
BCH	5.66–6.53 (6.10 ± 0.62)	NA	6.25–6.49 (6.37 ± 0.17)	6.64	NA	5.81	6.05	6.78	6.59–6.98	6.51–6.78 (6.62 ± 0.09)
ZYW	7.76–8.72 (8.24 ± 0.68)	NA	7.63–8.07 (7.85 ± 0.31)	7.38	7.85	NA	7.94	7.73	7.41–7.52	7.44–7.81 (7.69 ± 0.13)
MAW	7.03–7.04 (7.04 ± 0.01)	NA	6.13–6.68 (6.41 ± 0.39)	6.06	7.07	7.42	6.89	7.21	7.18–7.23	7.12–7.38 (7.31 ± 0.10)
IOW	3.51–4.39 (3.95 ± 0.62)	NA	3.98–4.40 (4.19 ± 0.30)	3.82	NA	4.09	4.31	3.84	3.67–3.78	3.68–3.86 (3.79 ± 0.06)
CM 3 L	4.46–5.31 (4.89 ± 0.60)	NA	4.32–4.44 (4.38 ± 0.08)	4.66	4.36	4.68	4.42	4.51	4.38–4.43	4.45–4.63 (4.53 ± 0.06)
C 1 C 1 W	2.19–2.45 (2.32 ± 0.18)	NA	2.21–3.32 (2.77 ± 0.78)	2.07	NA	NA	NA	3.22	3.01–3.21	3.06–3.28 (3.21 ± 0.08)
M 3 M 3 W	3.30–3.80 (3.55 ± 0.35)	NA	4.74–4.97 (4.86 ± 0.16)	3.38	4.36	4.88	4.83	4.74	4.59–4.61	4.62–4.74 (4.67 ± 0.04)
RCM	0.64–0.66 (0.65 ± 0.01)	NA	0.44–0.70 (0.57 ± 0.18)	0.61	0.73	0.69	0.67	0.68	NA	NA
CM 3 L	4.91–5.74 (5.33 ± 0.59)	NA	4.53–5.11 (4.82 ± 0.41)	5.08	4.36	5.13	4.09	5.03	4.71–4.81	4.88–5.03 (4.93 ± 0.05)
ML	8.85–9.22 (9.04 ± 0.26)	NA	8.77–9.06 (8.92 ± 0.21)	9.42	8.42	9.31	9.08	8.41	8.09–8.15	8.34–8.69 (8.49 ± 0.13)
MDL	8.72–9.49 (9.11 ± 0.54)	NA	9.24–9.46 (9.35 ± 0.16)	9.82	8.42	NA	NA	8.76	8.21–8.43	8.51–8.91 (8.75 ± 0.13)
CPH	3.30–3.61 (3.46 ± 0.22)	NA	2.99–3.20 (3.10 ± 0.15)	3.31	3.06	3.32	2.96	2.86	2.52–2.76	2.69–2.93 (2.83 ± 0.09)