Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi) Author Miranda, Gustavo Silva de 81150D94-592A-4CE5-8E88-E60F557A4341 Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark. & Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington DC, USA. smiranda.gustavo@gmail.com Author Giupponi, Alessandro P. L. 434112AC-B212-43E8-A5D9-2F5D5619AFC4 Laboratório de Referência Nacional em Vetores das Riquetsioses, LIRN-IOC-FIOCRUZ, Rio de Janeiro, RJ, Brazil. agiupponi@gmail.com Author Prendini, Lorenzo C2D080D0-75DB-4DA1-A101-AB4DCF50FF0A Arachnology Lab, Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA. lorenzo@amnh.org Author Scharff, Nikolaj F84D2235-66D2-460C-820D-80024068759D Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark. & Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington DC, USA. & Zoology Section, Natural History Museum of Denmark, University of Copenhagen, Denmark. nscharff@snm.ku.dk text European Journal of Taxonomy 2021 2021-09-24 772 1 409 http://dx.doi.org/10.5852/ejt.2021.772.1505 journal article 4042 10.5852/ejt.2021.772.1505 b65bb5a9-bbe7-49a4-af44-4b4c03121288 2118-9773 5536410 9B82A32F-0A07-47E3-8684-FED7C8EBF1E9

Table 4 (continued on next five pages). Measurements of all eastern South American species of Charinus Simon, 1892.

							Pedipalp								Leg IV
Species	Additional information	Source	Sex	Carapace length	Carapace width	Femur length	Patella length	Tibia length	Tarsus length	Claw length	Femur I length	Femur	Basitibia article I	Basitibia article II	Basitibia article III	Basitibia article IV	Distitibia	Basitarsus	Other tarsal articles
C. acaraje	MNRJ 9297	This work	prob.♂ no gonopod	3.60	5.19	4.00	3.84	1.88	1.25	1.04	12.56	7.28	3.56	1.25	1.31	1.64	3.80	2.00	1.20
C. acaraje	MNRJ 9297	This work	♀	2.25	3.72	2.45	2.38	1.25	0.80	0.80	8.30	5.38	2.34	0.75	0.88	1.15	2.66	1.50	0.98
C. acaraje	MNRJ 9297	This work	♀	2.25	4.20	3.00	2.92	1.50	0.94	0.94	10.90	6.06	–	–	–	–	–	–	–
C. acaraje	MNRJ 9297	This work	♀	3.06	4.38	3.08	2.92	1.25	0.94	1.00	10.77	6.06	2.80	0.94	1.00	1.30	3.06	1.70	1.00
C. acaraje	MNRJ 9297	This work	prob.♂ no gonopod	3.69	5.13	4.60	4.50	1.84	1.19	1.09	13.59	7.60	3.40	1.25	1.20	1.55	3.68	2.19	1.25
C. acaraje	MNRJ 9297	This work	prob.♂ no gonopod	3.88	5.38	4.44	4.38	1.97	1.25	1.25	13.46	8.20	3.90	1.35	1.45	1.88	4.10	2.16	1.13
C. acaraje	MNRJ 09125	This work	♀	2.60	3.65	2.20	2.08	1.04	0.80	0.76	7.52	5.00	2.24	0.80	0.68	1.00	2.60	1.41	0.84
C. alagoanus sp. nov.	MNRJ 9294	This work	♀	2.76	4.00	2.13	2.03	1.16	0.80	0.71	6.32	4.35	2.31	0.60	0.78	–	2.38	1.18	0.75
C. alagoanus sp. nov.	MNRJ 9295	This work	♀	2.80	3.76	2.44	2.40	1.25	0.91	3.70	7.36	4.75	2.63	0.69	0.88	–	2.63	1.25	0.83
C. apiaca sp. nov.	MNRJ 9286	This work	♀	4.13	5.75	4.50	4.50	1.88	1.35	1.20	12.82	7.60	3.80	0.92	1.40	1.96	3.44	1.43	1.08
C. asturius	MZSP 18930 holotype	Pinto-da-Rocha et al. 2002	♂	3.80	5.45	–	7.30	–	–	–	9.50	5.90	3.30	0.75	1	1.20	2.60	1.20	1.00
C. asturius	MZSP 18930 paratype	Pinto-da-Rocha et al. 2002	♀	2.60	3.60	–	2.30	–	–	–	5.35	3.60	1.75	0.45	0.5	0.70	1.75	0.75	0.75
C. asturius	MZSP 18934 female	Pinto-da-Rocha et al. 2002	♀	3.80	5.50	–	4.60	–	–	–	9.25	6.00	3.30	0.90	1.1	1.25	2.75	1.20	1.10
C. brasilianus	SMF 25397 paratype	This work	♀	3.00	4.50	2.55	2.45	1.15	0.84	0.72	6.80	–	–	–	–	–	–	–	–
C. brasilianus	SMF 25397 paratype	This work	♂	3.05	4.45	3.05	3.10	1.20	0.80	0.60		–	–	–	–	–	–	–	–
C. brasilianus	SMF 25397 paratype	This work	♂	3.68	5.28	7.50	7.50	1.60	0.88	0.90	7.95	–	–	–	–	–	–	–	–
C. brasilianus	SMF 25397 paratype	This work	♂	2.88	5.19	5.69	5.94	1.44	1.00	0.65	7.69	–	–	–	–	–	–	–	–
C. brasilianus	SMF 25397 paratype	This work	♂	3.44	5.52	6.67	6.79	1.80	1.10	0.80	8.97	–	–	–	–	–	–	–	–
C. brasilianus	MNRJ 9271 Santa Teresa	This work	♂	3.25	4.90	6.40	6.32	1.44	0.94	0.75	8.10	4.65	2.34	0.72	0.75	1.13	2.63	1.05	0.80
C. brasilianus	MNRJ 9271 Santa Teresa	This work	♀	3.32	5.00	2.96	3.08	1.44	0.93	0.82	7.84	5.00	–	–	–	–	–	–	–
Species	Additional information	Source	Sex	Carapace length	Carapace width	Femur length	Patella length	Tibia length	Tarsus length	Claw length	Femur I length	Femur	Basitibia article I	Basitibia article II	Basitibia article III	Basitibia article IV	Distitibia	Basitarsus	Other tarsal articles
C. brasilianus	MNRJ 9226 Aracruz	This work	♂	2.88	4.15	4.05	4.19	1.13	0.80	0.70	7.12	4.35	2.19	0.53	0.55	0.78	2.25	0.97	0.67
C. brasilianus	MNRJ 9226 Aracruz	This work	♂	2.88	4.15	4.05	4.19	1.13	0.80	0.70	7.12	4.35	2.19	0.53	0.55	0.78	2.25	0.97	0.67
C. brasilianus	MNRJ 9226 Aracruz	This work	♂	2.88	4.15	4.05	4.19	1.13	0.80	0.70	7.12	4.35	2.19	0.53	0.55	0.78	2.25	0.97	0.67
C. brasilianus	MNRJ 9226 Aracruz	This work	♀	2.80	4.20	2.40	2.44	1.13	0.82	0.68	7.20	4.70	2.25	0.60	0.64	0.82	2.50	1.12	0.72
C. brasilianus	MNRJ 9240 Aricanga	This work	subad. ♂	3.25	4.85	3.25	3.20	1.33	0.98	0.80	9.00	5.44	2.59	0.78	0.84	1.09	2.88	1.20	0.78
C. brasilianus	MNRJ 9240 Aricanga	This work	♀	2.80	4.15	2.22	2.25	1.10	0.80	0.69	6.80	4.45	2.13	0.52	0.60	0.75	2.35	1.00	0.70
C. brasilianus	MNRJ 9240 Aricanga	This work	♀	2.97	4.25	2.47	2.41	1.16	0.88	0.75	7.20	4.75	2.20	0.60	0.70	0.94	2.59	1.10	0.78
C. brasilianus	MNRJ 9240 Aricanga	This work	♂	3.10	4.38	4.38	4.56	1.38	0.86	0.70	8.20	4.90	2.34	0.68	0.75	0.93	2.76	1.15	0.75
C. brasilianus	MNRJ 9240 Aricanga	This work	♂	3.00	4.25	4.06	4.06	1.08	0.76	0.61	7.10	4.35	2.28	0.63	0.75	0.84	2.56	0.98	0.70
C. caatingae	Mean values	Vasconcelos & Ferreira 2016	♀	3.10	4.03	2.24	2.43	1.13	0.86	0.70	–	–	–	–	–	–	–	–	–
C. caatingae	Mean values	Vasconcelos & Ferreira 2016	♀	3.60	4.69	2.80	3.05	1.35	1.01	0.81	–	–	–	–	–	–	–	–	–
C. carinae sp. nov.	MNRJ 9293/9199	This work	♂	4.81	7.12	7.20	7.20	2.28	1.52	1.30	13.33	7.76	4.13	1.34	1.56	1.66	3.56	1.28	1.09
C. carinae sp. nov.	MNRJ 9293/9199	This work	♂	4.06	6.00	4.44	4.56	1.84	1.25	1.09	10.64	6.56	3.30	1.15	1.20	1.35	3.00	1.25	0.95
C. carioca sp. nov.	MNRJ 9201	This work	♀	4.69	6.80	5.44	5.31	2.28	1.84	1.38	18.72	10.51	6.08	1.90	2.25	2.15	4.15	1.78	1.38
C. carioca sp. nov.	MNRJ 9201	This work	♂	4.50	6.25	7.12	6.88	2.09	1.58	1.13	18.08	9.87	5.44	1.63	1.94	1.97	4.20	1.70	1.30
C. cearensis sp. nov.	IBSP 67	This work	♀	3.88	5.50	4.38	4.56	1.88	1.25	1.00	13.85	8.21	4.00	1.32	1.56	2.00	3.80	1.50	1.05
C. cearensis sp. nov.	CHNUFPI	This work	♀	3.35	4.60	3.10	3.25	1.47	0.90	0.84	10.51	5.88	3.00	1.09	1.16	1.34	3.40	1.22	0.84
C. diamantinus sp. nov.	1255 MNRJ 9203	This work	♀	4.50	6.13	3.50	3.55	1.72	1.60	1.08	11.28	6.8	3.75	1.24	1.28	1.64	3.36	1.22	1.06
C. diamantinus sp. nov.	MNRJ 9300	This work	♂	3.56	4.69	4.60	4.44	2.36	1.56	0.97	15.38	8.70	5.00	1.52	1.48	2.28	4.55	1.66	0.97
C. diamantinus sp. nov.	MNRJ 9300	This work	♂	3.60	4.70	3.85	3.60	1.88	1.28	1.03	14.10	7.92	4.10	1.40	1.44	2.00	4.00	1.56	0.80
C. diamantinus sp. nov.	MNRJ 9300	This work	juv. ♂	3.63	4.88	2.81	2.81	1.56	1.31	0.91	11.41	6.48	3.50	1.22	1.31	1.66	3.56	1.08	0.88
Species	Additional information	Source	Sex	Carapace length	Carapace width	Femur length	Patella length	Tibia length	Tarsus length	Claw length	Femur I length	Femur	Basitibia article I	Basitibia article II	Basitibia article III	Basitibia article IV	Distitibia	Basitarsus	Other tarsal articles
C. diamantinus sp. nov.	MNRJ 9299 paratype	This work	♀	5.68	7.76	5.92	5.76	3.08	2.52	1.50	17.95	10.90	6.16	2.35	2.40	2.85	5.00	1.70	1.40
C. diamantinus sp. nov.	MNRJ 9189	This work	♂	4.80	6.48	4.06	4.06	2.20	1.68	1.08	14.10	8.00	4.44	1.44	1.50	1.88	3.13	1.20	1.08
C. diamantinus sp. nov.	MNRJ 9190	This work	♂	5.60	7.52	5.84	5.88	2.63	2.24	1.25	16.54	9.20	3.19	1.75	1.88	2.44	4.19	1.48	1.28
C. eleonorae	MNRJ 9033	This work	♂	3.44	4.69	4.19	4.06	1.63	1.20	0.94	13.46	7.80	3.70	1.35	1.45	1.85	3.75	1.53	0.97
C. eleonorae	MNRJ 9033	This work	♀ with eggs	3.75	5.19	4.25	4.00	1.75	1.34	1.00	–	7.44	3.69	1.44	1.44	1.88	3.96	1.52	1.00
C. eleonorae	MNRJ 9033	This work	♀	3.44	4.80	3.75	3.75	1.25	1.25	0.85	12.05	6.80	3.31	1.32	1.40	1.60	3.60	1.44	0.94
C. eleonorae	MNRJ 9033	This work	♀	3.40	4.15	3.80	3.55	1.80	1.25	0.82	11.54	6.72	3.20	1.20	1.24	1.63	3.60	1.45	0.90
C. eleonorae	MNRJ 9033	This work		3.40	4.50	4.15	3.84	1.72	1.20	0.90	12.69	7.44	3.25	1.18	1.35	1.75	3.80	1.53	0.93
C. euclidesi sp. nov.	MNRJ 9099	This work	♂	4.63	6.48	9.50	9.30	2.16	1.52	1.11	15.51	8.60	4.25	1.41	1.56	2.03	4.25	1.60	1.25
C. euclidesi sp. nov.	MNRJ 9099	This work	♀	4.00	6.06	4.55	4.25	1.66	1.31	1.10	14.36	7.76	3.60	1.44	1.38	1.91	3.70	1.38	1.13
C. goitaca sp. nov.	MNRJ 9224 Vitória	This work	♂	4.75	6.88	7.60	7.60	2.09	1.50	1.13	15.00	8.50	4.65	1.31	1.47	2.13	4.30	1.38	1.06
C. goitaca sp. nov.	MNRJ Vila Velha	This work	subad. ♂	3.60	5.00	2.88	2.91	1.16	0.97	0.87	10.26	5.81	3.03	0.87	0.95	1.20	2.97	1.25	0.94
C. goitaca sp. nov.	MNRJ Vila Velha	This work	juv. ♀	2.88	4.00	2.00	1.94	1.00	0.81	0.73	7.36	4.60	–	–	–	–	–	–	–
C. goitaca sp. nov.	MNRJ Vila Velha	This work	juv. ♀	2.60	3.60	1.68	1.65	0.88	0.67	0.66	6.00	3.60	–	–	–	–	–	–	–
C. imperialis sp. nov.	MNRJ 9296	This work	♀	4.96	6.88	4.90	4.75	2.10	1.56	1.25	14.23	8.00	4.00	1.40	1.25	2.05	4.00	1.47	1.19
C. imperialis sp. nov.	MNRJ 9276	This work	juv. ♀	2.94	4.19	2.00	1.94	1.00	0.80	0.68	6.60	4.15	2.00	0.64	0.60	0.88	2.16	0.90	0.75
C. iuiu	Mean values	Vasconcelos & Ferreira 2016	?	3.22	4.46	2.50	2.79	1.34	1.01	0.68	–	–	–	–	–	–	–	–	–
C. iuiu	Mean values	Vasconcelos & Ferreira 2016	?	3.69	5.12	3.15	3.48	1.62	1.18	0.79	–	–	–	–	–	–	–	–	–
C. jibaossu	MNRJ 9217	This work	♀	4.00	6.10	4.56	4.38	1.75	1.38	1.10	11.92	7.04	3.56	1.18	1.20	1.50	3.48	1.25	0.93
C. jibaossu	MNRJ 9277	This work	♂	3.55	5.00	4.38	4.50	1.45	1.20	0.80	10.51	5.75	3.04	1.00	1.04	1.20	3.12	0.11	0.92
C. monasticus sp. nov.	MNRJ 9103 holotype	This work	♀	3.69	5.00	2.80	2.96	1.56	1.13	0.95	10.00	6.56	4.35	1.60	–	–	4.25	1.72	0.97
C. monasticus sp. nov.	MNRJ 9101	This work	♀	2.8	3.8	2.13	2.05	1.3	0.8	0.70	7.8	4.75	2.8	1.16	–	–	3.09	1.44	0.88
Species	Additional information	Source	Sex	Carapace length	Carapace width	Femur length	Patella length	Tibia length	Tarsus length	Claw length	Femur I length	Femur	Basitibia article I	Basitibia article II	Basitibia article III	Basitibia article IV	Distitibia	Basitarsus	Other tarsal articles
C. schirchii	Holotype	Mello-Leitão 1931	♀	4.00	5.00	4.00	5.00	–	–	–	–	–	–	–	–	–	–	–	–
C. sooretama sp. nov. C. sooretama sp. nov.	MNRJ 9245 MNRJ 9244	This work This work	♂ ♂	3.36 3.50	5.00 4.75	6.40 6.13	6.72 6.48	1.56 1.50	0.98 1.00	0.85 0.75	8.00 7.60	4.95 5.00	2.56 2.40	0.69 0.69	0.78 0.84	0.94 0.94	2.76 2.56	1.15 1.10	0.75 0.74
C. sooretama sp. nov.	MNRJ 9245	This work	♀	2.60	3.92	1.94	1.94	1.00	0.80	0.64	6.00	3.85	1.81	0.50	0.63	0.80	2.13	0.83	0.68
C. sooretama sp. nov.	MNRJ 9245	This work	♀	2.50	3.70	1.91	1.90	1.00	0.78	0.55	5.63	3.60	1.75	0.45	0.55	0.80	2.00	0.86	0.68
C. sooretama sp. nov.	MNRJ 9245	This work	♀	2.90	4.50	2.41	2.41	1.18	0.88	0.70	6.40	4.40	2.05	0.50	0.63	0.80	2.13	1.00	0.70
C. souzai sp. nov.	MNRJ 9090	This work	♀	2.88	4.50	2.64	2.64	1.25	0.90	0.73	6.96	5.00	2.19	0.56	0.63	0.94	2.41	0.97	0.71
C. souzai sp. nov.	MNRJ 9090	This work	♀	3.10	4.50	2.52	2.76	1.14	0.90	0.68	6.96	4.75	2.28	0.60	0.70	1.14	2.40	1.03	0.71
C. souzai sp. nov.	MNRJ 9090	This work	♀	2.95	4.35	2.40	2.36	1.05	0.80	0.71	6.64	4.40	2.85	–	–	–	2.41	0.96	0.70
C. souzai sp. nov.	MNRJ 9090	This work	♀	3.04	4.55	2.40	2.40	1.25	0.87	0.71	6.88	4.75	2.25	0.53	0.59	1.06	2.50	1.10	0.70
C. souzai sp. nov.	MNRJ 9090	This work	♀	2.75	4.25	2.53	2.50	1.22	0.84	0.71	6.40	4.81	2.22	0.55	0.68	1.03	2.45	0.98	0.66
C. souzai sp. nov.	MNRJ 9336	This work	♀	3.40	5.00	2.75	2.81	1.25	0.87	0.70	8.00	5.313	2.48	0.625	0.75	1.125	2.5	1.3	0.8
C. souzai sp. nov.	MNRJ 9336	This work	♀	2.95	4.40	2.69	2.50	1.23	0.92	0.70	7.36	4.75	2.13	0.60	0.63	1.08	2.75	1.02	0.66
C. spelaeus	Mean values	Vasconcelos & Ferreira 2017		2.3	3.03	2.16	2.46	1.00	0.75	0.63	–	–	–	–	–	–	–	–	–
C. taboa	Mean values	Vasconcelos et al. 2016	♀	3.21	4.63	3.11	3.3	1.34	0.96	0.74	–	–	–	–	–	–	–	–	–
C. taboa	Mean values	Vasconcelos et al. 2016	♂	3.31	4.69	4.56	5.02	1.43	1.03	0.72	–	–	–	–	–	–	–	–	–
C. troglobius	MNRJ 9069	This work	♂	4.00	5.04	4.56	4.56	2.34	1.91	1.13	23.33	11.54	6.40	2.50	2.88	3.00	4.81	2.20	1.36
C. troglobius	MNRJ 9069	This work	♀	3.35	4.25	2.88	2.92	1.60	1.36	0.84	12.56	6.56	3.75	1.48	1.40	1.72	3.16	1.25	0.95
C. una sp. nov.	MNRJ 9131	This work	♂	2.28	3.40	2.23	2.10	1.05	0.65	0.65	6.40	4.10	1.75	0.53	0.52	0.94	2.38	1.35	0.63
C. una sp. nov.	MNRJ 9131	This work	♀	2.08	3.00	1.53	1.43	0.80	0.60	0.52	4.69	3.40	1.30	0.35	0.38	0.73	1.93	1.13	0.62
C. una sp. nov.	MNRJ 9298	This work	♀	2.00	2.88	1.50	1.53	0.75	0.49	0.52	4.30	3.13	1.25	0.35	0.38	0.63	1.75	1.08	0.63
C. una sp. nov.	MNRJ 9132	This work	♀	2.08	3.00	1.60	1.58	0.82	0.60	0.58	5.00	3.60	1.08	0.40	0.43	0.75	2.00	1.18	0.70
Species	Additional information	Source	Sex	Carapace length	Carapace width	Femur length	Patella length	Tibia length	Tarsus length	Claw length	Femur I length	Femur	Basitibia article I	Basitibia article II	Basitibia article III	Basitibia article IV	Distitibia	Basitarsus	Other tarsal articles
C. madagascariensis	MNHN, syntype	This work	♂	2.60	3.80	2.03	2.03	0.94	0.69	0.75	5.00	2.97	1.90	0.56	0.48	0.38	1.19	0.56	0.56
C. madagascariensis	MNHN, syntype	This work	♀	2.80	4.00	2.19	2.18	1.13	0.72	0.83	5.28	3.40	1.50	0.40	0.46	0.56	1.78	0.68	0.66
C. madagascariensis	MNHN, syntype	This work	no gonopod	3.35	4.75	4.75	4.75	1.43	1.10	0.90	8.60	5.00	2.60	0.68	0.88	1.13	2.38	0.95	0.83
C. madagascariensis	MNHN, syntype	This work	no gonopod	3.32	5.00	3.85	3.80	1.38	1.08	0.85	9.20	5.56	–	–	–	–	–	–	–
C. madagascariensis	MNHN, syntype	This work	no body, legs only									–	2.59	0.88	0.94	1.16	2.53	1.06	0.88
C. madagascariensis	MNHN, syntype	This work	♀	2.90	4.35	2.92	2.88	1.25	0.86	0.81	6.72	4.40	2.09	0.50	0.68	0.83	2.03	0.88	0.75
C. madagascariensis	MNHN, syntype	This work	♀	3.65	5.44	4.50	4.60	1.66	1.18	0.87	9.49	5.63	2.92	0.83	1.00	1.25	2.75	1.04	0.80
C. madagascariensis	MNHN, syntype	This work	♀	3.44	4.90	3.80	4.00	1.56	1.00	1.06	7.44	4.90	2.50	0.68	0.70	0.88	2.50	1.00	0.80
C. susuwa sp. nov.	BMNH, holotype	This work	♀	3.13	4.88	3.92	3.88	1.85	1.20	0.90	11.41	7.36	4.15	1.09	1.38	1.53	4.10	1.88	1.13