Heth pivari n. sp. (Nematoda: Ransomnematoidea: Hethidae) from the indigenous North American millipede Narceus gordanus (Spirobolida: Spirobolidae), with keys for worldwide Heth spp. Author Phillips, Gary 0000-0003-1930-1009 Department of Entomology and Plant Pathology, University of Tennessee, 370 Plant Biotechnology Building, 2505 E J Chapman Drive, Knoxville, TN 37996 - 4560 USA. & https: // orcid. org / 0000 - 0003 - 1930 - 1009 Author Moulton, John K. 0000-0001-8760-3274 Department of Entomology and Plant Pathology, University of Tennessee, 370 Plant Biotechnology Building, 2505 E J Chapman Drive, Knoxville, TN 37996 - 4560 USA. & https: // orcid. org / 0000 - 0001 - 8760 - 3274 Author Bernard, Ernest C. 0000-0003-3359-1516 Department of Entomology and Plant Pathology, University of Tennessee, 370 Plant Biotechnology Building, 2505 E J Chapman Drive, Knoxville, TN 37996 - 4560 USA. & https: // orcid. org / 0000 - 0003 - 3359 - 1516 text Zootaxa 2020 2020-10-20 4861 4 486 514 journal article 9031 10.11646/zootaxa.4861.4.2 a15b3af7-16c7-492b-bf7b-d08a36b7f6a2 1175-5326 4416659 8B330C85-5B40-48EF-8C17-48332637C1C9

TABLE 4. Selected Heth male characteristics.

Species	Caudal papillae 2 & 4 close, medium or distant	Caudal alae present/ absent	Stoma wide or com- pressed	Cuticular ornamentationa	Sources other than original descrip- tion
H. adolphi	distant	absent	?	many lobes whole body length	―
H. albertoi	?	absent	?	spines in anterior region	―
H. amazonensis	?	absent	?	none	―
H. artigasi	close (Adamson)	absent	wide	none	Kloss (1965), Adamson (1983), Tantalean & Altmann (1988)
H. baracoa	close	absent	?	none	―
H. baudini	close	present	wide	none	―
H. bifidispiculum	close	absent	slightly compressed	none	―
H. clunyi	close	absent	slightly compressed	none	―
H. costata	no males	―	―	―	―
H. coyi	?	absent	?	none	―
H. dimorphum	?	absent	?	none	Kloss (1965)
H. duvidosum	no males	―	―	―	―
H. gordae	distant	absent	wide	none	―
H. hamatus	?	absent	?	none	―
H. hexaspinosum	?	absent	?	none	Kloss (1965)
H. hispaniolae	?	absent	?	spines in esophageal region	―
H. imias	close	absent	?	none	―
H. impalutiensis	close	absent	wide	none	―
H. initiensis	close	present	compressed	none
H. insularis	?	absent	?	none	―
H. josephinae	distant	absent	?	many elongated lobes whole body length	―
H. juli	?	?	?	none	Travassos (1929), Hunt (2015)
H. konoplevi	close	present	wide	none
H. lamothei	close	absent	?	none	―
H. macrocephala	?	absent	?	none	―
H. magnavulvaris	no males	―	―	―	―
H. maicuru	close (Adamson)	absent	?	none	Kloss (1965), Adamson (1983)