Heth pivari n. sp. (Nematoda: Ransomnematoidea: Hethidae) from the indigenous North American millipede Narceus gordanus (Spirobolida: Spirobolidae), with keys for worldwide Heth spp. Author Phillips, Gary 0000-0003-1930-1009 Department of Entomology and Plant Pathology, University of Tennessee, 370 Plant Biotechnology Building, 2505 E J Chapman Drive, Knoxville, TN 37996 - 4560 USA. & https: // orcid. org / 0000 - 0003 - 1930 - 1009 Author Moulton, John K. 0000-0001-8760-3274 Department of Entomology and Plant Pathology, University of Tennessee, 370 Plant Biotechnology Building, 2505 E J Chapman Drive, Knoxville, TN 37996 - 4560 USA. & https: // orcid. org / 0000 - 0001 - 8760 - 3274 Author Bernard, Ernest C. 0000-0003-3359-1516 Department of Entomology and Plant Pathology, University of Tennessee, 370 Plant Biotechnology Building, 2505 E J Chapman Drive, Knoxville, TN 37996 - 4560 USA. & https: // orcid. org / 0000 - 0003 - 3359 - 1516 text Zootaxa 2020 2020-10-20 4861 4 486 514 journal article 9031 10.11646/zootaxa.4861.4.2 a15b3af7-16c7-492b-bf7b-d08a36b7f6a2 1175-5326 4416659 8B330C85-5B40-48EF-8C17-48332637C1C9

TABLE 4. (Continued)

Species	Caudal papillae 2 & 4 close, medium or distant	Caudal alae present/ absent	Stoma wide or com- pressed	Cuticular ornamentationa	Sources other than original description
aAll males likely have somatic papillae. Cuticular ornamentation refers to conspicuous cuticular lobes or spines, not papillae.
H. mauriesi	close	absent	slightly compressed	none	Spiridonov & Yushin (2000), Carreno et al. (2013)
H. multiplus	?	absent	?	none	―
H. orthopori	close	absent	compressed	none	―
H. ortonwilliamsi	close	present	moderately com- pressed	none	―
H. parartigasi	no males	―	―	―	―
H. perarmatum	no males	―	―	―	―
H. pinnatum	no males	―	―	―	―
H. pivari n. sp.	close	absent	compressed	none	―
H. poeyi	?	absent	?	none	―
H. sinediscus	close	present?	?	none	―
H. spinalatum	?	absent	?	small spines in esophageal area	―
H. spinosum	close (Adamson); medium (T&A)	absent	slightly compressed	none	Adamson (1983), Tantalean & Altmann (1988)
H. spoliatus	?	absent	?	none	―
H. sutherlandi	no males	―	―	―	―
H. taybaci	medium	present	?	none	―
H. taynguyeni	medium	present	?	none	―
H. tonkinensis	medium	absent	wide	none	―
H. travassosi	?	absent	?	none	Kloss (1965)
H. travofilhoi	close (Adamson)	absent	moderately com- pressed	none	Adamson (1983)
H. tuxtlensis	close	absent	?	none	―
H. tuzetae	distant (Adamson)	absent	slightly compressed	none	Kloss (1965), Adamson (1983)
H. vietnamensis	distant	absent	?	none	―
H. xaniophora	close	present	moderately com- pressed	none	―
H. xarochae	distant	absent	?	none	―
H. zeuglocantha	no males	―	―	―	―