Integrative taxonomy reveals two new species of karst-dwelling Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from the border region of Laos and Vietnam Author Luu, Vinh Quang 0000-0002-0634-1338 Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam vinhlq@vnuf.edu.vn Author Hoang, Tuoi Thi 0000-0002-3564-0601 Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam tuoiht@vnuf.edu.vn Author Ha, Hong Bich 0000-0003-4777-5463 College of Forestry Biotechnology, Vietnam National University of Forestry, Hanoi, Vietnam honghb@vnuf.edu.vn Author Grismer, Jesse L. 0000-0002-2542-5430 jgrismer@lasierra.edu Author Murdoch, Matthew 0000-0001-5914-6408 hewmurdoch@gmail.com Author Sitthivong, Saly 0000-0002-9993-2136 Faculty of Forestry, National University of Laos, Dong Dok Campus, Vientiane, Lao PDR s.sitthivong@nuol.edu.la Author Phimpasone, Vilay 0009-0001-9744-0728 Faculty of Forestry, National University of Laos, Dong Dok Campus, Vientiane, Lao PDR v.phimpasone@nuol.edu.la Author Grismer, L. Lee 0000-0001-8422-3698 Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA & Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400, Kota Kinabalu, Sabah, Malaysia lgrismer@lasierra.edu text Zootaxa 2024 2024-07-24 5486 1 71 108 http://dx.doi.org/10.11646/zootaxa.5486.1.3 journal article 300860 10.11646/zootaxa.5486.1.3 120e528d-555c-43c7-beeb-fd4fa982267c 1175-5326 13209709 E696DDCA-FEAE-4EC6-9FF4-0CC3E5F55063

TABLE 4. Sex and raw meristic data used in the analyses from specimens of Hemiphyllodactylus members within clade 6 (excepting H. dushanensis Zhou and Liu, 1981 due to lack of individual specimens). m = male; f = female; sa=subadult.

Species	Museum no.	Sex				Meristic data
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			Chin	CN	SL	IL	VS	DS	SL1F	SL1T
banaensis	ITBCZ 2465	m	6	3	11	10	9	18	5	5
banaensis	ITBCZ 2461	m	6	3	11	9	10	18	5	5
banaensis	ITBCZ 2450	f	6	3	11	10	10	18	5	5
banaensis	ITBCZ 2462	f	7	3	12	11	11	20	5	5
banaensis	ITBCZ 2463	f	6	3	9	10	10	18	5	5
banaensis	ITBCZ 2464	f	6	3	10	10	10	17	5	5
banaensis	ITBCZ 2466	f	6	3	10	10	9	17	5	5
banaensis	ITBCZ 2467	f	6	3	12	11	10	19	5	4
banaensis	ITBCZ 2468	f	6	3	11	11	12	20	5	5
banaensis	ITBCZ 2469	f	7	3	11	11	10	20	5	5
bonkowskii	IEBR 4689	m	7	3	10	10	13	24.5	5	5
bonkowskii	IEBR 4690	m	7	3	9	9	15	25	5	5
bonkowskii	IEBR 4691	f	5	3	9	9	15	25	5	5
bonkowskii	IEBR 4693	f	7	3	9.5	8	15	25.5	5	4
bonkowskii	IEBR 4692	f	7	3	10	10.5	14	25	5	5
bonkowskii	IEBR 4749	saf	6	3	10	9	14	26	5	5
dupanglingensis	CSUFT 00401	f	10	4	11	12	11	15	5	5
dupanglingensis	CSUFT 00402	m	10	3	11	9	11	15	4	4
dupanglingensis	CSUFT 00403	m	9	4	11	9	10	16	5	4
dupanglingensis	CSUFT 00404	m	11	3	12	10	10	15	5	5
dupanglingensis	CSUFT 00405	m	11	3	11	10	11	14	5	5
dupanglingensis	CSUFT 00406	m	11	3	14	9	11	14	5	5
hongkongensis	SYS r001735	m	5	3	10	9.5	10	15	5	5
			Chin	CN	SL	IL	VS	DS	SL1F	SL1T
hongkongensis	SYS r001728	f	6	3	11	10	10	13	4	5
hongkongensis	SYS r001729	f	5	3	12	10	9	14	4	5
hongkongensis	SYS r001730	f	6	3	12	10	9	13	4	5
hongkongensis	SYS r001731	f	6	3	10	10	9	12	3	5
hongkongensis	SYS r001732	f	5	4	11.5	10.5	9	14	4	5
hongkongensis	SYS r001733	f	6	3	11	10.5	10	13	4	5
hongkongensis	SYS r001734	m	6	3	11	10.5	9	14	5	5
houaphanensis sp. nov.	VNUF R.2020.17	f	8	2	12	8	10	18	3	4
houaphanensis sp. nov.	VNUF R.2020.16	m	9	2	12	11	11	18	4	4
houaphanensis sp. nov.	NUOL R.2020.20	m	8	2	11	9	13	23	4	4
huishuiensis	NJNUh00851	m	10	3	11	11	9	14	3	3
huishuiensis	NJNUh00859	m	8	3	11	10.5	9	15	3	3
huishuiensis	NJNUh00852	f	10	3	10	10	8	13	3	3
huishuiensis	NJNUh00854	f	10	3	9	9	7	13	3	3
huishuiensis	NJNUh00855	f	10	3	10	9	8	13	3	3
huishuiensis	NJNUh00856	f	10	3	10	10	7	13	3	3
huishuiensis	NJNUh00857	f	8	3	10	10	8	14	3	3
huishuiensis	NJNUh00858	f	10	3	10	10	9	14	3	3
kiziriani	IEBR A.2014.3	m	9	4	11	10.5	13.5	21.5	5	5
kiziriani	IEBR A.2014.4	m	9	4	11	9	14.5	26	5	5
kiziriani	VNMN A.2014.1	m	7	4	10	10	13.5	22.5	5	5
kiziriani	ZFMK 95702	m	6	4	10.5	9	14	23.5	5	5
kiziriani	IEBR A.2014.5	f	6	4	10.5	10	11.5	20.5	5	5
			Chin	CN	SL	IL	VS	DS	SL1F	SL1T
kiziriani	NUOL R - 2014.1	f	9	4	11	10.5	13.5	23.5	5	5
kiziriani	NUOL R - 2014.2	f	6	4	10	10	11	18	5	5
kiziriani	VNMN A.2014.2	f	8	4	11	9	12	24	5	5
kiziriani	ZFMK 95703	f	8	4	10	10.5	13	25	5	5
kiziriani	ZFMK 95704	f	9	4	10.5	9.5	12.5	23.5	5	5
lungcuensis	VNUF R.2021.01	m	8	3	12	10.5	7	13	4	4
lungcuensis	IEBR	m	8	3	11	11	6	13	4	4
lungcuensis	VNUF R.2021.03	m	9	3	12	10	7	15	4	4
lungcuensis	IEBR	f	9	2	11	10	9	16	4	4
lungcuensis	VNUF R.2021.08	f	10	3	11	10	7	12	3	4
lungcuensis	VNUF R.2023.01	m	10	3	11.5	9.5	10	17	4	4
lungcuensis	VNUF R.2023.02	f	10	3	11	9.5	11	16	4	4
nahangensis	IEBR 4741	m	8	3	11	9.5	9	22	3	3
nahangensis	IEBR 4742	m	8	2.5	11.5	10.5	9	18	3	3
nahangensis	IEBR 4743	f	9	3	11	10.5	13	23	3	3
ngocsonensis	IEBR 4694	m	8	3	10.5	10	14	20.5	5	5
ngocsonensis	IEBR 4695	m	6	3	10	10	13	20.5	4	5
ngocsonensis	IEBR 4696	f	8	3	10	9	15	19.5	5	5
ngocsonensis	IEBR 4697	f	8	3	10	9.5	14	19.5	5	6
serpispecus	NUOL 00476	m	7	3	11	9	10	26	4	4
vanhoensis sp. nov.	VNUF R.2022.09	f	9	5.5	11.5	10.5	10	15	4.5	5
vanhoensis sp. nov.	VNUF R.2022.07	f	7	5	11	11	8	19	4	4
vanhoensis sp. nov.	VNUF R.2023.22	j	10	5	10	10.5	11	18	5	5
typus	JAV bg. R.39	f	10	6	10.5	10	9	14	4	4
			Chin	CN	SL	IL	VS	DS	SL1F	SL1T
typus	MS92002	f	15	5	12	11	12	15	4	4
typus	CUMZ R 2003.19	f	11	5	11.5	11	11	15	4	4
typus	RBWS 19309	f	15	5	11.5	11	12	16	4	5
yanshanensis	KIZ 062090	m	9	6	9.5	10	8	14	5	5
yanshanensis	KIZ 062091	f	8	5	11.5	11	7	15	5	5
yanshanensis	KIZ 062092	f	11	5	11	10.5	8	15	5	5
yanshanensis	KIZ 062093	f	8	5	10.5	10	13	22	5	5
yanshanensis	KIZ 062094	f	9	5	12	9.5	7	14	5	5
yanshanensis	KIZ 062095	f	8	5	11	10.5	7	13	5	6
yanshanensis	KIZ 062096	f	8	5	11.5	11	8	15	4	5
yanshanensis	KIZ 062097	m	8	5	10.5	10	7	14	4	5
yanshanensis	KIZ 062098	m	9	5	10	9.5	8	18	5	5
yanshanensis	KIZ 062099	m	8	5	11	11	8	15	5	5
yanshanensis	KIZ 062100	f	8	5	10	9.5	11	20	5	5
yanshanensis	KIZ 062101	f	10	5	11	10	8	14	4	5
yanshanensis	KIZ 062102	f	10	5	10.5	9.5	7	11	4	5
zugi	IEBR A.2013.20	m	9	3	12.5	11	15	21.5	4	4.5
zugi	ZFMK 94781	m	12	2	11	10.5	15	20.5	5	5
zugi	IEBR A.2013.21	m	9	2.5	10.5	11	16	21.5	5	5