Integrative taxonomy reveals two new species of karst-dwelling Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from the border region of Laos and Vietnam Author Luu, Vinh Quang 0000-0002-0634-1338 Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam vinhlq@vnuf.edu.vn Author Hoang, Tuoi Thi 0000-0002-3564-0601 Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam tuoiht@vnuf.edu.vn Author Ha, Hong Bich 0000-0003-4777-5463 College of Forestry Biotechnology, Vietnam National University of Forestry, Hanoi, Vietnam honghb@vnuf.edu.vn Author Grismer, Jesse L. 0000-0002-2542-5430 jgrismer@lasierra.edu Author Murdoch, Matthew 0000-0001-5914-6408 hewmurdoch@gmail.com Author Sitthivong, Saly 0000-0002-9993-2136 Faculty of Forestry, National University of Laos, Dong Dok Campus, Vientiane, Lao PDR s.sitthivong@nuol.edu.la Author Phimpasone, Vilay 0009-0001-9744-0728 Faculty of Forestry, National University of Laos, Dong Dok Campus, Vientiane, Lao PDR v.phimpasone@nuol.edu.la Author Grismer, L. Lee 0000-0001-8422-3698 Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA & Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400, Kota Kinabalu, Sabah, Malaysia lgrismer@lasierra.edu text Zootaxa 2024 2024-07-24 5486 1 71 108 http://dx.doi.org/10.11646/zootaxa.5486.1.3 journal article 300860 10.11646/zootaxa.5486.1.3 120e528d-555c-43c7-beeb-fd4fa982267c 1175-5326 13209709 E696DDCA-FEAE-4EC6-9FF4-0CC3E5F55063

TABLE 5. Significant p - values from the results of the TukeyHSD tests analyses comparing the normalized morphometric and meristic characters of Hemiphyllodactylus houaphanensis sp. nov. to other species of the Indochina lineage within clade 6. Abbreviations are in the Materials and methods. * indicates which characters used a Welch’s ANOVA.

Morphometric characters	SVL	Trunk	HeadL	HeadW	SnEye	NarEye	EyeD	SnW	Chin	CN	SL	VS	DS	SL1F	SL1T
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houaphanensis – banaensis	0.003				0.004	<0.001				<0.001
houaphanensis – bonkowskii		<0.001		<0.001	<0.001	<0.001		<0.001		<0.001	0.007		0.004	0.004	0.004
huishuiensis – houaphanensis		<0.001		0.023						<0.001		0.024	<0.001	<0.001	<0.001
lungcuensis – houaphanensis		<0.001				0.011	<0.001			0.002		0.031	0.009	0.009
nahangensis – houaphanensis		<0.001								0.027
serpispecus – houaphanensis		0.005
vanhoensis – houaphanensis		<0.001
zugi – houaphanensis		<0.001			<0.001
kiziriani – houaphanensis			0							<0.001
typus– houaphanensis			<0.001	<0.001	<0.001	0.024	<0.001	<0.001	<0.001	0
yanshanensis – houaphanensis			<0.001	0.002		0.017				0		0.018	0.030	0.030	0.030
ngocsonensis – houaphanensis				<0.001	<0.001	<0.001				<0.001
houaphanensis – dupanglingensis					0.002	0.014			0.026	<0.001			0.024	0.024	0.024
houaphanensis – hongkongensis										<0.001			<0.001	<0.001	<0.001
vanhoensis – houaphanensis										0