Species richness under a vertebral stripe: integrative taxonomy uncovers three additional species of Pholidobolus lizards (Sauria, Squamata, Gymnophthalmidae) from the north-western Colombian Andes Author Amezquita, Adolfo Laboratory of Biodiversity and Cloud Forests Conservation, Bioconservancy. org, Jardin, Colombia aamezquita@bioconservancy.org Author Mazariegos-H, Luis A. https://orcid.org/0000-0003-4882-0414 Laboratory of Biodiversity and Cloud Forests Conservation, Bioconservancy. org, Jardin, Colombia Author Canaveral, Santiago Laboratory of Biodiversity and Cloud Forests Conservation, Bioconservancy. org, Jardin, Colombia Author Orejuela, Catalina Department of Biological Sciences, Universidad de los Andes, Bogota, Colombia Author Barragan-Contreras, Leidy Alejandra Department of Biological Sciences, Universidad de los Andes, Bogota, Colombia Author Daza, Juan M. Grupo Herpetologico de Antioquia, Universidad de Antioquia, Medellin, Colombia jumadaza@gmail.com text ZooKeys 2023 2023-01-19 1141 119 148 http://dx.doi.org/10.3897/zookeys.1141.94774 journal article http://dx.doi.org/10.3897/zookeys.1141.94774 1313-2970-1141-119 55DA386251E649CAA4CFF4534D96A267 A1754208B2555011B8DFE90160CF9343

Table 3. Meristic and morphometric traits. Summary of meristic and morphometric (in mm) traits in adult lizards of the four clades of Pholidobolus studied here. Mean +/- sd (min - max).

Trait	P. argosi sp. nov. (n = 21)	P. celsiae sp. nov. (n = 11)	P. marianus comb. nov. (n = 24)	P. odinsae sp. nov. (n = 35)
Prefrontals	0 +/- 0 (0-0)	2.0 +/- 0.0 (2-2)	2.0 +/- 0.0 (2-2)	2.0 +/- 0.0 (2-2)
Supraoculars	2.0 +/- 0.0 (2-2)	3.0 +/- 0.0 (3-3)	3.4 +/- 0.5 (3-4)	3.0 +/- 0.2 (3-4)
Superciliaries	3.9 +/- 0.4 (3-5)	3.9 +/- 0.3 (3-4)	3.7 +/- 0.6 (3-5)	4.0 +/- 0.2 (3-5)
Lower palpebrals	3.9 +/- 0.7 (2-5)	4.4 +/- 0.7 (3-5)	5.4 +/- 1.1 (4-7)	3.9 +/- 0.8 (2-5)
Suboculars	4.1 +/- 0.6 (3-5)	4.9 +/- 0.9 (4-6)	3.9 +/- 0.8 (3-6)	5.2 +/- 1.1 (3-7)
Postoculars	2.0 +/- 0.0 (2-2)	2.1 +/- 0.3 (2-3)	2.9 +/- 0.6 (2-4)	2.2 +/- 0.5 (2-4)
Temporal	12.5 +/- 1.8 (9-17)	21.8 +/- 4.8 (14-28)	13.8 +/- 2.7 (10-19)	19.9 +/- 3.3 (11-26)
Supralabials	6.9 +/- 0.4 (6-8)	7.2 +/- 0.4 (7-8)	6.9 +/- 0.5 (6-8)	7.1 +/- 0.4 (7-8)
Infralabials	5.6 +/- 0.6 (5-7)	4.6 +/- 0.7 (4-6)	4.9 +/- 0.7 (3-6)	4.8 +/- 0.7 (4-6)
Pregulars	2.1 +/- 0.2 (2-3)	2.4 +/- 0.5 (2-3)	3.8 +/- 0.9 (2-6)	2.3 +/- 0.6 (2-4)
Gulars	7.1 +/- 0.4 (6-8)	7.6 +/- 0.5 (7-8)	7.9 +/- 1.3 (4-10)	8.1 +/- 0.6 (7-9)
Collar scales	10.3 +/- 1.5 (7-13)	11.2 +/- 1.6 (9-14)	10.3 +/- 1.7 (6-13)	10.9 +/- 2.0 (6-17)
Dorsal transverse	30.1 +/- 1.0 (28-32)	29.5 +/- 0.9 (28-31)	30.3 +/- 1.2 (28-32)	29.9 +/- 1.0 (28-32)
Dorsal longitudinal	18.8 +/- 1.2 (17-21)	24.9 +/- 1.0 (23-26)	22.3 +/- 1.9 (19-26)	22.5 +/- 2.0 (20-26)
Around mid-body	31.3 +/- 2.3 (26-35)	39.6 +/- 1.5 (37-43)	35.9 +/- 2.6 (30-42)	38.4 +/- 3.4 (31-45)
Transversal ventral	21.1 +/- 0.7 (20-22)	19.6 +/- 1.0 (18-21)	21.2 +/- 1.2 (19-24)	19.8 +/- 1.2 (17-23)
Head width	8.70 +/- 1.20 (6.6-11.3)	11.89 +/- 1.20 (7.4-14.6)	7.53 +/- 0.98 (5.9-9.7)	8.02 +/- 1.02 (6.0-10.0)
Head length	11.77 +/- 1.79 (9.1-15.0)	15.05 +/- 2.46 (10.4-17.9)	10.66 +/- 1.56 (7.6-14.5)	11.24 +/- 1.39 (8.6-15.0)
Head height	6.13 +/- 0.87 (5.0-7.7)	8.44 +/- 1.51 (5.2-9.7)	5.32 +/- 0.71 (3.9-7.1)	5.41 +/- 0.63 (4.2-6.7)
Jaw length	9.51 +/- 1.42 (7.2-13.2)	12.77 +/- 1.56 (9.4-14.5)	10.15 +/- 1.88 (7.1-14.1)	9.67 +/- 1.50 (7.4-13.2)
Longest finger	5.25 +/- 0.58 (4.1-6.4)	5.80 +/- 0.79 (4.7-6.8)	4.75 +/- 0.47 (4.0-5.6)	4.72 +/- 0.63 (3.9-6.3)
Pelvis width	6.57 +/- 0.70 (4.8-7.6)	8.60 +/- 1.01 (6.2-9.6)	6.57 +/- 0.91 (4.2-7.9)	6.67 +/- 0.97 (4.9-8.9)
Longest toe	8.13 +/- 0.77 (6.8-10.2)	9.42 +/- 0.90 (7.4-10.2)	7.19 +/- 0.65 (5.7-8.2)	7.19 +/- 1.06 (5.3-10.8)
Tail width	5.50 +/- 0.59 (4.3-6.6)	7.61 +/- 1.42 (5.1-9.5)	4.97 +/- 0.60 (4.1-6.1)	5.69 +/- 0.97 (3.9-8.1)
Snout-vent length	50.85 +/- 4.23 (42.6-57.9)	62.38 +/- 7.19 (45.0-68.6)	46.97 +/- 4.81 (35.6-55.8)	48.36 +/- 5.71 (35.4-60.3)