Leaving no stone unturned: three additional new species of Atractus ground snakes (Serpentes, Colubridae) from Ecuador discovered using a biogeographical approach Author Arteaga, Alejandro https://orcid.org/0000-0002-0014-3728 Biodiversity Field Lab (BioFL), Khamai Foundation, Quito, Ecuador af.arteaga.navarro@gmail.com Author Quezada, Amanda https://orcid.org/0000-0001-9121-2583 Biodiversity Field Lab (BioFL), Khamai Foundation, Quito, Ecuador Author Vieira, Jose https://orcid.org/0000-0003-0926-0362 ExSitu, Quito, Ecuador Author Guayasamin, Juan M. Laboratorio de Biologia Evolutiva, Instituto Biosfera, Colegio de Ciencias Biologicas y Ambientales COCIBA, Universidad San Francisco de Quito USFQ, Quito, Ecuador text ZooKeys 2022 2022-09-15 1121 175 210 http://dx.doi.org/10.3897/zookeys.1121.89539 journal article http://dx.doi.org/10.3897/zookeys.1121.89539 1313-2970-1121-175 15C9C77E688849BBBC57F7500EF2E06C 5ED52281A14D529BAB575B813B1D599D
Table 3. Differences in coloration, scale counts, and size between Atractus arangoi and A. major. The range of each continuous variable is from our own sample, Prado (1939), and Maynard et al. (2017). The numbers in parentheses represent the sample size.
Variable characterAtractus arangoiAtractus major
Dark brown or black nape stripeAbsentPresent
Dorsal markingsIrregular dark blotchesComplete irregular dark bands anteriorly; blotches posteriorly
SexMales (n = 2)Females (n = 2)Males (n = 7)Females (n = 5)
Maximum SVL309 mm412 mm533 mm986 mm
Ventral scales154-163160-161162-165172-177
Subcaudal scales38-3929-3236-4534-37