Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data Author Kavanaugh, David H. Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA dkavanaugh@calacademy.org Author Maddison, David R. https://orcid.org/0000-0002-7152-3824 Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA Author Simison, W. Brian Center for Comparative Genomics, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA Author Schoville, Sean D. https://orcid.org/0000-0001-7364-434X Department of Entomology, University of Wisconsin, Madison, WI 53706, USA Author Schmidt, Joachim Institute of Biosciences, University of Rostock, Universitaetsplatz 2, D- 18055 Rostock, Germany Author Faille, Arnaud Department of Entomology, Coleoptera, Stuttgart State Museum of Natural History, Rosenstein 1, 70191 Stuttgart, Germany Author Moore, Wendy https://orcid.org/0000-0003-4441-6203 Department of Entomology, University of Arizona, Tucson, AZ 85721 - 0036, USA Author Pflug, James M. Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA Author Archambeault, Sophie L. Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Berkeley, 142 Weill Hall # 3200, Berkeley, CA 94720, USA Author Hoang, Tinya Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA Author Chen, Jei-Ying Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Santa Cruz, Long Marine Lab, 117 McAllister Way, Santa Cruz, CA 95060, USA text ZooKeys 2021 2021-06-16 1044 41 152 http://dx.doi.org/10.3897/zookeys.1044.62245 journal article http://dx.doi.org/10.3897/zookeys.1044.62245 1313-2970-1044-41 C6AE8C5BC5D64A09A26E00FAADBF86E1 535A79F3B89F5699A71E2B83B65C4C21

. For each specimen, DNA voucher number is listed under "DNA ID" and its depository is listed under "Dep" (AF = A. Faille collection; CAS = California Academy of Sciences; JS = J. Schmidt collection; KS = K. Sasakawa collection; OSAC = Oregon State University Arthropod Collection; SDS = S.D. Schoville collection).

Taxon	DNA ID	Locality	Dep
Leistus birmanicus	DHK0084	China, Yunnan, Gongshan County, Dulongjiang Township, Bapo, Miliwang, 1956 m, 27.72383°N, 098.36117°E	CAS
Leistus crenatus	DHK1939	Algeria, Central Djurdjura, Anou Boussouil, 1718 m, 36.4690°N, 4.1914°E	CAS
Leistus ferrugineus	DHK1903	Italy, Piedmont, Cuneo, Po River Valley at Paesano, 618 m, 44.67584°N, 7.27709°E	CAS
Leistus ferruginosus	DHK0336	USA, Oregon, Benton County, Alsea Falls, 326 m, 44.31949°N, 123.49750°W	CAS
Leistus frateroides	SDS13-123A	Russia, Altai Republic, Evrechala Mt., 1891 m, 51.47490°N, 087.42992°E	CAS
Leistus fulvibarbis	DHK1937	France, Eure, La Saussaye, 49.25326°N, 0.99201°E	CAS
Leistus fulvus	DHK1938	Azerbaijan, Qax, forest litter in road to Qax, 580 m, 41.39828°N, 46.92153°E	CAS
Leistus gaoligongensis	DHK0340	China, Yunnan, Longyang County, Luoshuidong, 2308 m, 24.94833°N, 98.75667°E	CAS
Leistus kryzhanovskii	SDS13-330A	Russia, Altai Republic, Aigulaksky Range, 2555 m, 50.44896°N, 087.40025°E	CAS
Leistus lihengae	DHK0343	China, Yunnan, Longyang County, Luoshuidong, 2308 m, 24.94833°N, 98.75667°E	CAS
Leistus longipennis	DHK0338	USA, California, Del Norte County, 8.3 miles N of Crescent City, 18 m, 41.86559°N, 124.13906°W	CAS
Leistus madmeridianus	DHK0339	USA, California, Sonoma County, Salt Point State Park, 91 m, 38.56823°N, 123.31916°W	CAS
Leistus niger	DHK1761	Russia, Republic of Khakassia, Karasibo River, 750 m, 52.30713°N, 90.12761°E	CAS
Leistus niitakaensis	DHK1800	Taiwan, Chiayi County, Alishan District, west slope Yu Shan, 3429 m, 23.46641°N, 120.95041°E	CAS
Leistus nitidus	DHK1888	France, Hautes-Alpes, Ristolas, 1643 m, 44.77196°N, 6.95399°E	CAS
Leistus nokoensis	DHK1834	Taiwan, Tiachung City, Xueshan, Black Forest, 3340 m, 24.39359°N, 121.24936°E	CAS
Leistus nubivagus	DHK0502	Canary Islands, Tenerife, La Victoria, Barranco del Madrono, 1435 m	CAS
Leistus parvicollis	DHK2107	Greece, Kefallonia, Enos, 1000-1600 m	CAS
Leistus pyrenaeus	DHK1941	Spain, Lleida, Llastarri, Minas de Canal	CAS
Leistus rufomarginatus	DHK1394	Belgium, West Flanders, Wijnendale Forest	CAS
Leistus sardous	DHK1942	Italy, Sicily, Piana degli Albanesi - Grotta del Garrone	AF
Leistus smetanai	DHK1860	Taiwan, Taichung City, Xueshan, Black Forest, 3460 m, 24.39373°N, 121.25114°E	CAS
Leistus sp JIL 2	SDS12-202	China, Jilin, northern slope of Changbai Shan, 1772 m, 42.04913°N, 128.05782°E	CAS
Leistus sp NEP	DHK2108	Nepal, Lapchi Kang range, Chera, 4400 m, 27.89861°N, 86.06278°E	CAS
Leistus sp SCH	DHK0634	China, Sichuan, Litang County, Shalulishan, Haizishan Yakou, 4610 m, 29.47366°N, 100.21921°E	CAS
Leistus sp YUN 1	DHK0015	China, Yunnan, Gongshan County, Heiwadi, 2070 m, 27.79650°N, 98.58267°E	CAS
Leistus sp YUN 2	DHK0224	China, Yunnan, Shibali Yakou, second cirque, 3700 m, 27.20333°N, 98.69303°E	CAS
Leistus taiwanensis	DHK1838	Taiwan, Nantou County, Tatajia at Dongpu Cabin, 2563 m, 23.48438°N, 120.88591°E	CAS
Nebria acuta	DHK0018	USA, Alaska, Wrangell Mts., Kennicott, National Creek, 615 m, 61.48469°N, 142.88696°W	CAS
Nebria aenea	DHK1445	Russia, Altai Krai, Charyshsky District, Altai Mts., Gorelyi, Korgon River, 1000 m, 53.0°N, 83.8°E	CAS
Nebria albimontis	SDS05-018	USA, California, Mono County, North Fork Perry Eiken Creek, 3278 m, 37.63879°N, 118.22910°W	CAS
Nebria altaica	DHK0404a	Russia, Buryat Republic, Tunka Mts., East Branch Kingarga River, 1500 m, 51.97275°N, 102.45970°E	CAS
Nebria angustata	DHK1653	Switzerland, Bern, Guttannen, Furtwangsattel	CAS
Nebria angusticollis	DHK1896	France, Hautes-Alpes, Col d'Agnel, 2704 m, 44.68499°N, 6.97789°E	CAS
Nebria appalachia	DHK0507	USA, Tennessee, Sevier County, Great Smoky Mountains National Park, Ramsey Prong, 1084 m, 35.7088°N, 83.3129°W	CAS
Nebria arinae	SDS12-063E	Russia, Altai Republic, Holzun Mts., 2162 m, 50.20821N °, 084.55408°E	CAS
Nebria arkansana	DHK0838	USA, Colorado, Summit County, Quandary Peak, 3763 m, 39.39159°N, 106.11371°W	CAS
Nebria austriaca	DHK1654	Austria, Lower Austria, The Oscher	CAS
Nebria baenningeri	SDS12-025A	Russia, Altai Republic, Holzun Mts., 2018 m, 50.31269°N, 084.61154°E	CAS
Nebria baicalica	DHK0386a	Russia, Irkutsk Oblast, Lake Baikal at Bolschie Koty, 450 m, 51.90405°N, 105.08014°E	CAS
Nebria balli	DHK0930	USA, Washington, Mt. Rainier National Park, Paradise area, Edith Creek Basin, 1807 m, 46.79678°N, 121.72711°W	CAS
Nebria banksii	DHK0406	Russia, Kamchatka, Avancha River, 11 m, 53.18297°N, 158.39328°E	CAS
Nebria baumanni	DHK1814A	USA, Nevada, Clark County, Spring Mts., Deer Creek, 2570 m, 36.31018°N, 115.62334°W	CAS
Nebria bellorum	DHK0510	USA, Tennessee, Sevier County, Great Smoky Mountains National Park, Middle Fork Little Pigeon River, 630 m, 35.7031°N, 83.3581°W	CAS
Nebria beverlianna	DHK0804	USA, Wyoming, Sublette County, Hoback River, 1930 m, 43.27364°N, 110.52606°W	CAS
Nebria boschi	DHK1509	Germany, Baden-Wuettemberg, Tuebingen, Bad Urach, Hoehle, 670 m, 48.480°N, 9.368°E	CAS
Nebria bremii	DHK1648	Switzerland, Uri, Schlossbergluecke	CAS
Nebria brevicollis OR	DHK0717	USA, Oregon, Polk County, 1.5 miles W of Dallas, 160 m, 44.90748°N, 123.35466°W	CAS
Nebria brevicollis SP	DHK0009	Spain, Albacete, Penascosa, 38.67305°N, 2.40324°E	CAS
Nebria businskyorum	DHK0676	China, Xizang, Medog, Baibung, E of Doxong Pass, 3937 m, 29.48748°N, 94.95791°E	CAS
Nebria calva	DHK0430a	USA, Arizona, Apache County, West Fork Little Colorado River, 2814 m, 33.95821°N, 109.51614°W	CAS
Nebria capillosa	DHK2029	Nepal, Dolpo district, Rupghat Khola valley E of Juphal, 2150-2500 m, 29.97°N, 82.85°E	CAS
Nebria carbonaria	DHK0459a	Russia, Kuril Islands, Paramushir, inland from Severo-Kurilsk, east slope Ebeko Volcano, 50.67533°N, 156.09550°E	CAS
Nebria carri	DHK0208	USA, Idaho, Camas County, Carrie Creek, 2426 m, 43.59555°N, 114.69264°W	CAS
Nebria cascadensis	DHK0899	USA, Washington, Whatcom County, North Fork Nooksack River, 222 m, 48.90471°N, 121.99183°W	CAS
Nebria castanea	DHK1519	Italy, Trentino-Alto, Passo di Gavia, 2547 m, 46.35983°N, 10.50054°E	CAS
Nebria castanipes	DHK0001	Canada, British Columbia, Alexander Creek, 3.6 miles W of Crowsnest Pass, 1308 m, 49.65580°N, 114.73222°W	CAS
Nebria catenata	DHK1067	USA, Utah, Grand County, La Sal Mts., Mill Creek, 2545 m, 38.50914°N, 109.28248°W	CAS
Nebria catenulata	DHK0405a	Russia, Buryat Republic, Tunka Mts., East Branch Kingarga River, 1500 m, 51.97275°N, 102.45970°E	CAS
Nebria changaica	DHK1451	Russia, Altai Republic, Altai Mts., Ukok Upland near Murzdy-Bulak Lake, 2700-3100 m, 49.28°N, 87.65°E	CAS
Nebria charlottae	DHK1254	Canada, British Columbia, Haida Gwaii, Graham Island, north shore at Tow Hill, 3 m, 54.07382°N, 131.66662°W	CAS
Nebria chinensis	DHK0600	China, Shaanxi, Zhouzhi County, Houzhenzi village, 1271 m, 33.85190°N, 107.83776°E	CAS
Nebria chuskae	DHK0437a	USA, Arizona, Apache County, Lukachukai Creek, 2200 m, 36.44443°N, 109.17993°W	CAS
Nebria coloradensis	DHK0852	USA, Colorado, Lake County, Lake Creek, 2927 m, 39.06532°N, 106.42093°W	CAS
Nebria complanata	DHK1949	Italy, Grosseto, Parco Regionale della Maremma, Monti dell'Uccellina, 42.63284°N, 11.07733°E	CAS
Nebria coreica	DRM4721	Russia: Primorsky Krai, 20 km W of Nachodka	CAS
Nebria crassicornis	DHK0021	USA, Washington, Olympic National Park, Hurricane Ridge, 1580 m, 47.97166°N, 123.49000°W	CAS
Nebria dabanensis	DHK0402b	Russia, Buryat Republic, Khamar-Daban Mts., Baikalsky Preserve, 1803 m, 51.34774°N, 105.13042°E	CAS
Nebria danmanni	DHK1134	USA, Washington, Olympic National Park, Mount Olympus, Snow Dome, 2050 m, 47.81165°N, 123.70196°W	CAS
Nebria darlingtoni	DHK0023	USA, California, El Dorado County, South Fork American River, 934 m, 38.76730°N, 120.48415°W	CAS
Nebria delicata	DHK2082	China, Xizang, Mt. Mendju Zari SE of Lhasa, 5200-5420m, 29.57°N, 91.22°E	JS
Nebria cf. desgodinsi	DHK2026	Nepal, Solu Khumbu district, near Kothe (Mosom Kharka), Inkhu Khola, 3600-3700 m, 27.67°N, 86.83°E	CAS
Nebria desolata	DHK0450a	USA, Utah, Garfield County, 6.0 miles NE of Henrieville, Dry Creek, 1971 m, 36.44443°N, 109.17993°W	CAS
Nebria diaphana	DHK1652	Italy, Trentino, Passo di Brocon	CAS
Nebria diversa	DHK0014	USA, Oregon, Lincoln County, 2 miles N of Newport at Moolack Beach, 6 m, 44.70572°N, 124.06115°W	CAS
Nebria dubatolovi	SDS12-092A	Russia, Altai Republic, Katunskii Mts., 2419 m, 50.00761°N, 086.25175°E	CAS
Nebria edwardsi	DHK0025	USA, Montana, Mineral County, Trout Creek, 13.4 miles NE of Hoodoo Pass, 1006 m, 47.07933°N, 114.92120°W	CAS
Nebria edwardsi	DHK0463a	USA, Montana, Mineral County, Trout Creek, 13.4 miles NE of Hoodoo Pass, 1007 m, 47.07933°N, 114.92120°W	CAS
Nebria eschscholtzii	DHK0022	USA, California, Somona County, Mark West Creek, 142 m, 38.54291°N, 122.72066°W	CAS
Nebria fontinalis	DHK1536	Switzerland, Uri, Oberalp, 2036 m, 46.66236°N, 8.66794°E	CAS
Nebria formosana	DHK1592	Taiwan, Taichung City, Heiping District, Xue Shan, Black Forest, 3460 m, 24.39372°N, 121.25114°E	CAS
Nebria fragariae	DHK0996	USA, Oregon, Grant County, Strawberry Creek, 1750 m, 44.31951°N, 118.67473°W	CAS
Nebria fragilis	DHK0028	USA, Utah, Utah County, South Fork American Fork River, 2207 m, 40.43519°N, 111.63603°W	CAS
Nebria frigida	DHK0005	USA, Alaska, Wrangell Mts., Kennicott, National Creek, 615 m, 61.48469°N, 142.88696°W	CAS
Nebria fulgida	DHK0409a	Russia, Buryat Republic, Tunka Mts., Kingarga River, 1380 m, 51.97478°N, 102.41099°E	CAS
Nebria fulviventris	DHK1397	Italy, Tuscany, Vallombrosa, 43.733°N, 11.557°E	CAS
Nebria gagates	DHK1880	France, Hautes-Alpes, Ristolas, Torrent de Clot Lucette, 1740 m, 44.76780°N, 6.94974°E	CAS
Nebria gebleri	DHK0002	USA, Alaska, Chichagof Island, Goulding River, 23 m, 57.77752°N, 136.25931°W	CAS
Nebria georgei	DHK0501	USA, Arizona, Grand Canyon National Park, Colorado River Mile 180, 36.187°N, 113.106°W	CAS
Nebria germari AU	DHK1645	Austria, Upper Austria, Hoellengebirge, Hoellkogel	CAS
Nebria germari GE	DHK1646	Germany, Bavaria, Allgaeuer Alps, Ifen	CAS
Nebria giulianii	SDS06-031B	USA, California, Inyo County, Milner Creek, 2241 m, 37.59480°N, 118.29763°W	CAS
Nebria gouleti	DHK0006	USA, Idaho, Idaho County, Selway River, 7.5 miles SE of Lowell, 475 m, 46.08804°N, 115.50410°W	CAS
Nebria gregaria	DHK0017	USA, Alaska, Aleutian Islands, Chuginadak Island, north shore, 10 m, 52.88447°N, 169.74653°W	CAS
Nebria gyllenhali	DHK0010	Russia, Buryat Republic, Khamar-Daban Mts., Tankhoy, 512 m, 51.53806°N, 105.112049°E	CAS
Nebria haida	DHK1300	Canada, British Columbia, Haida Gwaii, Moresby Island, ridge N of Takakia Lake, 837 m, 52.93787°N, 132.04862°W	CAS
Nebria holzunensis	SDS12-033D	Russia, Altai Republic, Holzun Mts., 1981 m, 50.25637°N, 084.59192°E	CAS
Nebria hudsonica	DHK0381a	USA, Wyoming, Sublette County, Hoback River, 1956 m, 43.24509°N, 110.48779°W	CAS
Nebria ingens	SDS08-025G	USA, California, Inyo County, North Fork Big Pine Creek, 3446 m, 37.11192°N, 118.51174°W	CAS
Nebria intermedia OR	DHK0990	USA, Oregon, Baker County, Elkhorn Range, Anthony Lake, 2182 m, 44.96122°N, 118.23200°W	CAS
Nebria intermedia UT	DHK1082	USA, Utah, Sevier County, 8.2 miles SE of Monroe, 2707 m, 38.55808°N, 112.08167°W	CAS
Nebria intermedia WY	DHK0809	USA, Wyoming, Teton County, Togwotee Pass, 2910 m, 43.75231°N, 110.06836°W	CAS
Nebria japonica	DHK1371	Japan, Honshu, Miyagi Prefecture, Road 12 E of Mt. Zao, 1462 m,	CAS
Nebria jeffreyi	DHK0039	USA, Oregon, Harney County, Steens Mts., Little Blitzen River, 2663 m, 42.69593°N, 118.59060°W	CAS
Nebria jockischi	DHK1526	Switzerland, Valais, Talgletscher Glacier, 2316 m, 46.45237°N, 7.83339°E	CAS
Nebria kagmara	DHK2120	Nepal, Karnali, Dolpo District, Kagmara Lekh below Kagmara La, 4400 m	CAS
Nebria kazabi	DHK1795	Russia, Republic of Tuva, Western Tannu-Ola Range, 2625 m, 50.54529°N, 90.99348°E	CAS
Nebria kincaidi	DHK0874	USA, Washington, Olympic National Park, Grand Pass, 1750 m, 47.86979°N, 123.35916°W	CAS
Nebria komarovi	DRM5171	China [locality unspecified]	CAS
Nebria korgonica	DHK1475	Russia, Altai Krai, Charyshsky District, Altai Mts., Gorelyi, Belok Mt., headwaters of Sentelek River, 1900 m, 51.075°N, 83.583°E	CAS
Nebria labontei	DHK0984	USA, Oregon, Wallowa County, West Fork Wallowa River below Glacier Lake, 2479 m, 45.16528°N, 117.28163°W	CAS
Nebria lacustris	DHK0004	USA, Maryland, Montgomery County, Potomac River at Plummers Island, 17 m, 38.96967°N, 77.18517°W	CAS
Nebria cf. laevistriata	DHK0492	China, Yunnan, Gongshan County, Bingzhongluo Township, 0.9 km N of Chukuai Lake, 4050 m, 27.99075°N, 098.47512°E	CAS
Nebria lafresnayei	DHK1946	Spain, Huesca, Villanua, Sinistro (Gouffre El)	CAS
Nebria lamarckensis	SDS06-648F	USA, California, Inyo County, side canyon of Pine Creek, 2350 m, 37.37605°N, 118.68400°W	CAS
Nebria lassenensis	DHK0220	USA, California, Shasta County, Lake Helen, 2485 m, 40.4689°N, 121.5131°W	CAS
Nebria laticollis	DHK1529	France, Haute-Savoie, Vallorcine, NW of Refuge Loriaz, 1962 m, 46.03847°N, 6.90832°E	CAS
Nebria lewisi	DHK1378	Japan, Honshu, Tochigi Prefecture, Fujioka, Watarase-sitch	CAS
Nebria ligurica	DHK1889	France, Hautes Alpes, 5.5 km SE of St. Veran, 2640 m, 44.66354°N, 6.92478°E	CAS
Nebria limbigera	DHK2083	China, Xinjiang, south slope of Wusunshan (= Ketmen) Mts., NNE of Zhaosu, 3080 m, 43.37667°N, 81.23861°E	CAS
Nebria lindrothi	DHK0020	USA, Colorado, Mineral County, Wolf Creek, 2 miles W of Wolf Creek Pass, 3180 m, 37.47989°N, 106.78024°W	CAS
Nebria lituyae	DHK0016	USA, Alaska, Juneau area, Mount Roberts Trail above treeline, 718 m, 58.29450°N, 134.37492°W	CAS
Nebria livida	DHK0007	Russia, Irkutsk Region, Tibielti River at Tunka Valley highway, 664 m, 51.76619°N, 103.24932°E	CAS
Nebria lombarda	DHK1400	Italy, Lombardia, Val di Scalve	CAS
Nebria louiseae	DHK1278	Canada, British Columbia, Haida Gwaii, Talunkwan Island, north shore, 2 m, 57.45150°N, 131.66662°W	CAS
Nebria lyelli	DHK0054	USA, California, Mono County, below Conness Lakes, 3134 m, 37.97600°N, 119.29690°W	CAS
Nebria lyubechanskii	SDS13-286A	Russia, Republic of Tuva, Western Tannu-Ola Range, north slope, 2744 m, 50.54470°N, 91.01261°E	CAS
Nebria macra	DHK1968	China, Xizang, Nyainqentanglha Shan, N Yangpachem, Bhuda valley, 5000-5200 m, 30.18222°N, 90.48917°E	CAS
Nebria macrogona	DHK1377	Japan, Honshu, Yamagata Prefecture, Kaminoyama, Sen'nin-sawa, Mts. Zao	CAS
Nebria mannerheimii	DHK0026	USA, Idaho, Idaho County, Salmon River at Riggins, 685 m, 45.32450°N, 116.34933°W	CAS
Nebria maroccana	DHK1945	Morocco, Fez-Meknes, Ifane, Ain Leuh to Azrou, Ifri Mkhrouga	CAS
Nebria martensi	DHK2069	Nepal, Jaljale Himal, below Paanch Pokhari, 4000-4100 m, 27.49028°N, 87.46111°E	JS
Nebria meanyi CA	DHK1357	USA, California, Siskiyou County, north slope of Mt. Shasta at Bolam Creek, 1850 m, 41.47177°N, 122.23174°W	CAS
Nebria meanyi WA	DHK0949	USA, Washington, Mt. Rainier National Park, Van Trump Creek above Christine Falls, 1164 m, 46.78242°N, 121.78021°W	CAS
Nebria mellyi	DHK1752	Russia, Kemerovskaya Oblast, Mustag Mt., 1290 m, 53.02153°N, 87.95143°E	CAS
Nebria mentoincisa	DHK2017	China, Xizang, Gangdise Shan, Kurum Valley NW Lhasa, SW Namba side valley, 4800-5150 m, 29.67528°N, 90.75444°E	CAS
Nebria metallica	DHK0915	USA, Washington, Pierce County, White River at Silver Creek Campground, 802 m, 46.99584°N, 121.53311°W	CAS
Nebria modoc	DHK0544	USA, California, Modoc County, Warner Mts., Thoms Creek, 1925 m, 41.56274°N, 120.27076°W	CAS
Nebria morvani	DHK0065	Nepal, Karnali, Jumla District, 2 km W of Guthichau, 2800 m, 29.20028°N, 82.30139°E	CAS
Nebria murzini	DHK0378a	China, Xinjiang, Tian Shan, Usu, 1500 m	CAS
Nebria nana	DHK1952	China, Tibet, S Tanggula Pass, 5000 m, 32.68500°N, 91.87278°E	CAS
Nebria navajo	DHK0451a	USA, Arizona, Navajo County, Keet Seel Creek, 1980-2040 m, 36.73423°N, 110.50803°W to 36.75269°N, 110.49601°W	CAS
Nebria niitakana	DHK1595	Taiwan, Taichung City, Heiping District, Xue Shan, Black Forest, 3460 m, 24.39372°N, 121.25114°E	CAS
Nebria niohozana	DHK1372	Japan, Honshu, Gunma Prefecture,Road 466 W of Shirane-san, 1883 m	CAS
Nebria nivalis AK	DHK0388a	USA, Alaska, Katmai National Park, Brooks Lake, 23 m, 58.54625°N, 58.54625°W	CAS
Nebria nivalis NU	DHK0387a	Canada, Nunavut, Baffin Island, Glasgow Inlet, Kimmirut, 40 m, 62.84728°N, 69.87410°W	CAS
Nebria nivalis gr sp	DHK1802	Russia, Republic of Tuva, Mongun-Taiga, 2490 m, 50.27584°N, 89.93906°E	CAS
Nebria nudicollis	DHK1944	Algeria, Central Djurdjura, Takouatz Guerissene, Tijdja a Tizi N'Kouilal	AF
Nebria numburica	DHK2075	Nepal, Solu Khumbu district, S of Dudh Kund, 4400-4600 m, 27.70°N, 86.58333°E	CAS
Nebria obliqua CO	DHK0436a	USA, Colorado, La Plata County, Durango, Animas River, 1971 m, 37.25852°N, 107.87664°W	CAS
Nebria obliqua UT	DHK1104	USA, Utah, Iron County, Parowan Creek, 2.0 miles S of Parowan, 1965 m, 37.81081°N, 112.80845°W	CAS
Nebria ochotica	DHK0403a	Russia, Kuril Islands, Paramushir, inland from Krasheninnikova Bay, Krasheninnikova River, 50.28517°N, 155.35650°E	CAS
Nebria ohdaiensis	DHK1380	Japan, Honshu Nara Prefecture, Kamikitayama, Mts. Ohdaigahara	CAS
Nebria olivieri	DHK2046	France, Ariege, Foret des Hares, Etang de Laurenti, 1650-2100 m, 42.68°N, 2.03°E	CAS
Nebria oowah	DHK1071	USA, Utah, Grand County, La Sal Mts., Oowah Lake, 2678 m, 38.50171°N, 109.27390°W	CAS
Nebria ovipennis	DHK0019	USA, California, Tuolumne County, Blue Canyon, 2800 m, 38.315°N, 119.659°W	CAS
Nebria oxyptera	DRM5172	China, Xinjiang Uyghur Autonomous Region, Pishan County, Xiadulla	CAS
Nebria pallipes	DHK0011	USA, Maryland, Montgomery County, Potomac River at Plummers Island, 17 m, 38.96967°N, 77.18517°W	CAS
Nebria paradisi OR	DHK1349	USA, Oregon, Hood River County, Mt. Hood Meadows Ski Area, 1890-2000 m, 45.34272°N, 121.67799°W	CAS
Nebria paradisi WA	DHK0929	USA, Washington, Mt. Rainier National Park, Paradise area, Edith Creek Basin, 1807 m, 46.79678°N, 121.72711°W	CAS
Nebria pasquineli	DHK0831	USA, Colorado, Boulder County, Lefthand Creek, 2420 m, 40.06590°N, 105.45711°W	CAS
Nebria pektusanica	SDS12-197E	China, Jilin, northern slope of Changbai Shan, 1901 m, 42.03954°N, 128.05562°E	CAS
Nebria perlonga	DHK0029	China, Xinjiang, Tian Shan, Usu, 1500 m	CAS
Nebria pertinax	DHK2119	Nepal, Mahakali/Darchula near Rapla, Shipu Lekh, 4300 m	JS
Nebria picea	DHK1649	Switzerland, St. Gallen, Saentis-Lisengrat	CAS
Nebria picicornis	DHK1876	France, Hautes Alpes, Aiguilles, Le Guil Torrent, 1454 m, 44.77790°N, 6.85961°E	CAS
Nebria piperi	DHK0965	USA, Washington, Mt. Rainier National Park, Nisqually River, W of Longmire, 774 m, 46.73469°N, 121.83055°W	CAS
Nebria piute	DHK1098	USA, Utah, Beaver County, North Fork Three Creeks, 3044 m, 38.32708°N, 112.37001°W	CAS
Nebria praedicta	SDS08-423b	USA, California, Trinity County, Thompson Peak, upper Grizzly Lake Basin, 2411 m, 41.00458°N, 123.04799°W	CAS
Nebria prezwalskii	DHK1950	China, Qinghai, S of Mt. Maqen Gangri, 4360 m, 34.55028°N, 98.14667°E	CAS
Nebria psammodes	DHK1908	France, Vaucluse, Toulourenc River at Brantes, 473 m, 44.19050°N, 5.33387°E	CAS
Nebria pseudorestias	DHK2081	Nepal, Taplejung distr., S of Kangchenjunga Himal, Timbu(wa) Pokhari, 4350-4500 m, 27.43°N, 88.05°E	CAS
Nebria purpurata CO	DHK0843	USA, Colorado, Summit County, Quandary Peak, 3763 m, 39.39159°N, 106.11371°W	CAS
Nebria purpurata NM	DHK1034	USA, New Mexico, Taos County, Red River at Junebug Campground, 2620 m, 36.70716°N, 105.43314°W	CAS
Nebria quileute	DHK0456a	USA, Washington, Olympic National Park, Boulder Creek, 47.97607°N, 123.69291°W	CAS
Nebria rathvoni	DHK0734	USA, California, Tuolumne County, Deadman Creek, 2667 m, 38.31763°N, 119.66534°W	CAS
Nebria retingensis	DHK2070	China, Xizang, Gangdise Shan, side valley S of Reting, 4500-4800 m, 30.26583°N, 91.53444°E	CAS
Nebria riversi	SDS08-377G	USA, California, Mono County, lake due S of Donohue Pass, 3377 m, 37.75289°N, 119.24847°W	CAS
Nebria roborowskii	DHK1955	China, Qinghai, pass Bayankala, 4700-4800 m, 34.12694°N, 97.65722°E	CAS
Nebria roddi	SDS13-086A	Russia, Altai Republic, north slope Krasnaya Mt., 2013 m, 50.07248°N, 085.22120°E	CAS
Nebria rubripes	DHK1871	France, Cantal, Le Lioran, 1212 m, 45.07996°N, 2.74542°E	CAS
Nebria rubrofemorata	DHK1464	Russia, Altai Krai, Altai Mts., watershed of Belogolosov Korgon and Inya Rivers, 2000-2250 m, 50.95°N, 83.645°E	CAS
Nebria saeviens	DHK1376	Japan, Honshu, Niigata Prefecture, Road 403 NE of Nozawaonsen, 841 m	CAS
Nebria sahlbergii	DHK0968	USA, Washington, Mt. Rainier National Park, Nisqually River, W of Longmire, 774 m, 46.73469°N, 121.83055°W	CAS
Nebria sajanica	DHK0401b	Russia, Buryat Republic, Tunka Mts., Kingarga River, 1852 m, 51.98045°N, 102.37566°E	CAS
Nebria sajanica gr sp 1	DHK1452	Russia, Republic of Tuva, Western Sayan Mts., near Kozhagar Mt., 2250-2450 m, 51.54°N, 89.17°E	CAS
Nebria sajanica gr sp 1	DHK1453	Russia, Republic of Tuva, Western Sayan Mts., near Kozhagar Mt., 2250-2450 m, 51.54°N, 89.17°E	CAS
Nebria sajanica gr sp 2	DHK1783	Russia, Krasnoyarsky Krai, Western Sayan Mts., Oiskiji Pass, 1385 m, 52.87277°N, 93.25745°E	CAS
Nebria sarlyk	DHK1735	Russia, Altai Republic, Sarlyk Mt., 2040 m, 51.061050°N, 85.690640°E	CAS
Nebria sayana	SDS13-205A	Russia, Republic of Khakassia, Samblyl Mt., headwaters of Karasibo River, 1890 m, 52.15164°N, 090.18011°E	CAS
Nebria schwarzi	DHK0427	Canada, Alberta, Cline River at Highway 11, 1325 m, 52.16983°N, 116.48304°W	CAS
Nebria sevieri	DHK1111	USA, Utah, Iron County, Parowan Creek, 12.5 miles S of Parowan, 2785 m, 37.71458°N, 112.84760°W	CAS
Nebria shiretokoana	SDS12-263A	Japan, Hokkaido, Shiretoko Peninsula, Rausu-daira, 1234 m, 44.07454°N, 145.13095°E	CAS
Nebria sierrablancae	DHK0431a	USA, New Mexico, Lincoln County, Sierra Blanca Ski Area, 3000 m, 33.39998°N, 105.78932°W	CAS
Nebria sierrae	DHK0052	USA, California, Mono County, below Conness Lakes, 3134 m, 37.97600°N, 119.29690°W	CAS
Nebria siskiyouensis	DHK0423a	USA, California, Trinity County, Big Flat Campground, 1490 m, 41.06465°N, 122.93493°W	CAS
Nebria sitnikovi	SDS12-131A	Russia, Altai Republic, Bashchelakskii Mts., 2012 m, 51.24219°N, 84.20432°E	CAS
Nebria snowi	DHK0460a	Russia, Kuril Islands, Ushishir Archipelago, Yankicha Island, inland from Kraternaya Bay, 20 m, 47.50867°N, 152.81950°E	CAS
Nebria sonorae	DHK0785	USA, California, Tuolumne County, Blue Canyon Creek, 2822 m, 38.31068°N, 119.66147°W	CAS
Nebria sp GAN	DHK2068	China Gansu, Lianhuashan, Shaltan, 2850 m	CAS
Nebria sp MG 1	DHK1138	Mongolia, Bayankhongor Province, northern slope of Ikh Bogd Uul, 1730 m, 44.979°N, 100.638°E	CAS
Nebria sp MG 2	DHK1139a	Mongolia, Oemnoegovi Province, Gobi Gurvanshaikhan National Park, Yolyn Am, 2285 m, 43.496°N, 104.089°E	CAS
Nebria sp XIZ 14	DHK0684	China, Xizang, Mainling, Paiqu, W of Doxong Pass, 4060 m, 29.48978°N, 094.93735°E	CAS
Nebria sp XIZ 16	DHK0679	China, Xizang, Bomi Zhamo, Garlungla, 3975 m, 29.76340°N, 95.70164°E	CAS
Nebria sp XIZ 17	DHK0681	China, Xizang, Bomi Zhamo, Garlungla, 3975 m, 29.76340°N, 95.70164°E	CAS
Nebria sp XIZ 20	DHK2008	China, Xizang, Transhimalaya, south slope Lungmari Mts., 4900-5300 m, 30.45°N, 86.50°E	CAS
Nebria sp XIZ 5	DHK0682	China, Xizang, Medog, Baibung, E of Doxong Pass, 4074 m, 29.49009°N, 94.95566°E	CAS
Nebria sp YUN 1	DHK0013	China, Yunnan, Tengchong County, Wuhe Township, Xiaodifang village, 2150 m, 24.43300°N, 098.75000°E	CAS
Nebria sp YUN 4	DHK0489a	China, Yunnan, Gongshan County, Bingzhongluo Township, 0.8 km N of Chukuai Lake, 3950 m, 27.98817°N, 098.47436°E	CAS
Nebria sp YUN 6	DHK0003	China, Yunnan, Longling Couty, Zhen'an Township, Bangbie village, 1540 m, 24.81306°N, 098.81667°E	CAS
Nebria sp YUN 9	DHK0087	China, Yunnan, Gongshan County, Cikai, Nu Jiang at Dashaba, 1443 m, 27.73606°N, 98.67113°E	CAS
Nebria sp YUN 10-dark	DHK0024	China, Yunnan, Tengchong County, Jietou Township, Yongangiao, 1500 m, 25.32556°N, 98.60000°E	CAS
Nebria sp YUN 10-pale	DHK0366b	China, Yunnan, Tengchong County, 2 km E of Qushi, 25.23944°N, 098.61667°E	CAS
Nebria sp YUN 11	DHK0086	China, Yunnan, Gongshan County, 0.6 km N of Dizhengdang, Dulong, 1898 m, 28.08680°N, 98.32835°E	CAS
Nebria sp YUN 13	DHK0483a	China, Yunnan, Gongshan County, Cikai Township, 0.1 km SE of Heipu Yakou, 3720 m, 27.76978°N, 098.44681°E	CAS
Nebria sp YUN 18	DHK0651	China, Yunnan, Lijiang County, Laojunshan, 3659 m, 26.64237°N, 99.74392°E	CAS
Nebria spatulata	SDS06-227B	USA, California, Fresno County, Sphinx Lakes, upper lake below Sphinx Crest, 3385 m, 36.71427°N, 118.51487°W	CAS
Nebria splendida	DHK2085	China, Xinjiang, W of Boro-Horo Mts., Mynmaral Mts., 1880 m, 44.10083°N, 82.19889°E	CAS
Nebria steensensis	DHK0042	USA, Oregon, Harney County, Steens Mts., Little Blitzen River, 2663 m, 42.69593°N, 118.59060°W	CAS
Nebria subdilatata	DHK0012	Russia, Buryat Republic, Khamar-Daban Mts., Tankhoy, 503 m, 51.53636°N, 105.10694°E	CAS
Nebria cf. superna	DHK1979	China, Xizang, Transhimalaya, pass Gyatso La S of Lhatse, 5200-5300 m, 28.95278°N, 87.43750°E	CAS
Nebria suturalis AB	DHK0435	Canada, Alberta, toe of Athabasca Glacier, 2047 m, 52.20819°N, 117.23597°W	CAS
Nebria suturalis CO	DHK0837	USA, Colorado, Clear Creek County, Mt. Evans, 4312 m, 39.58790°N, 105.64229°W	CAS
Nebria suturalis NH	DHK1151	USA, New Hampshire, Coos County, Mt. Washington, 1886 m, 44.26909°N, 71.30353°W	CAS
Nebria sylvatica	DHK0415a	USA, Washington, Olympic National Park, Boulder Creek, 655 m, 47.97607°N, 123.69291°W	CAS
Nebria tekesensis	DHK2084	China, Xinjiang, Narat Mts., W of Sarytur Pass, 3280 m, 42.90417°N, 82.62139°E	CAS
Nebria teletskiana	SDS12-153C	Russia, Altai Republic, Simultinskii Mts., Yambash Mt., 2006 m, 51.36033°N, 087.13060°E	CAS
Nebria tibialis	DHK1395	Italy, Tuscany, Vallombrosa, 43.733°N, 11.557°E	CAS
Nebria triad	DHK0471a	USA, California, Trinity County, South Fork Salmon River at Big Flat Campground, 1500 m, 41.06465°N, 122.93493°W	CAS
Nebria trifaria	DHK0434a	USA, Utah, Utah County, South Fork American Fork River, 2207 m, 40.43519°N, 111.63603°W	CAS
Nebria turcica	DHK1947	Turkey, Erzincan, Sipikor Gecidi, 2433 m, 39.88472°N, 39.56507°E	CAS
Nebria turmaduodecima	SDS08-424a	USA, California, Trinity County, Thompson Peak, upper Grizzly Lake Basin, 2411 m, 41.00458°N, 123.04799°W	CAS
Nebria uenoiana	DHK1588	Taiwan, Chiayi County, AlishanTownship, Laonong River, 3020 m, 23.40131°N, 121.93831°E	CAS
Nebria utahensis	DHK1075	USA, Utah, Garfield County, Henry Mountains, Lonesome Beaver, 2510 m, 38.10945°N, 110.77838°W	CAS
Nebria vandykei	DHK0932	USA, Washington, Mt. Rainier National Park, Paradise area, Edith Creek Basin, 1807 m, 46.79678°N, 121.72711°W	CAS
Nebria wallowae	DHK0980	USA, Oregon, Wallowa County, West Fork Wallowa River below Glacier Lake, 2479 m, 45.16528°N, 117.28163°W	CAS
Nebria walteriana	DHK1959	China, Xizang, NE of pass Shogu La, 5000-5350 m, 29.91333°N, 90.14111°E to 29.95556°N, 90.13028°E	CAS
Nebria wyeast	DHK0418b	USA, Oregon, Clackamas County, Mt. Hood, Timberline Lodge, 1876 m, 45.33539°N, 121.70849°W	CAS
Nebria yuae	DHK0603	China, Sichuan, Luding County, Gongga Shan, 3753 m, 29.58493°N, 102.02437°E	CAS
Nebria yunnana	DHK0377a	China, Yunnan, Zhanyi Co., Huashan Reservoir, 2000 m, 25.75°N, 103.61°E	CAS
Nebria zioni	DHK1108	USA, Utah, Iron County, Parowan Creek, 12.5 miles S of Parowan, 2785 m, 37.71458°N, 112.84760°W	CAS
Nippononebria altisierrae	DHK1547	USA, California, Alpine County, Carson Pass, 2616 m, 38.69377°N, 119.98749°W	CAS
Nippononebria campbelli	SDS13-388A	USA, Washington, Mt. Rainier National Park, Edith Creek Falls, 1613 m, 46.79051°N, 121.73016°W	CAS
Nippononebria chalceola	SDS14-629	Japan, Honshu, Fukui Prefecture, Ikeda Town, Mt. Heko, 1300 m	CAS
Nippononebria changbaiensis	SDS12-199a	China, Jilin, northern slope of Changbai Shan, 1901 m, 42.03954°N, 128.05562°E	CAS
Nippononebria virescens	DHK0348	USA, Oregon, Benton County, 0.3-0.5 miles W of Alsea Falls, 346 m, 44.32065°N, 123.50012°W	CAS
Notiophilus borealis	DHK0273b	USA, Alaska, Denali National Park, Wonder Lake, 731 m, 63.46222°N, 150.83167°W	CAS
Notiophilus semistriatus	DHK1335	USA, Idaho, Shoshone County, near Hobo Cedar Grove, 1425 m, 47.08477°N, 116.11538°W	CAS
Notiophilus sierranus	DHK1490	USA, California, El Dorado County, Blodgett Experimental Forest, 1300 m, 38.90958°N, 120.66123°W	CAS
Notiophilus sylvaticus	DHK1292	Canada, British Columbia, Haida Gwaii, Graham Island, Chinukundl Creek, 7 m, 53.32412°N, 131.95519°W	CAS
Opisthius richardsoni	DHK0885	USA, Washington, Whatcom County, Nooksack River at Cedarville, 45 m, 48.84072°N, 122.29334°W	CAS
Paropisthius chinensis	DHK0612	China, Sichuan, Luding County, Gongga Shan, 3045 m, 29.57393°N, 101.99204°E	CAS
Paropisthius davidis	DHK0645	China, Yunnan, Lijiang County, Laojunshan, 3502 m, 26.64210°N, 99.76745°E	CAS
Paropisthius indicus	DHK0329a	China, Yunnan, Cikae Township, Dabadi,3022 m, 27.79655°N, 98.50562°E	CAS
Paropisthius masuzoi	DHK1583	Taiwan, Nantou County, Ren'ai Township, Hehuanshan, 3090 m, 24.14367°N, 121.28075°E	CAS
Pelophila borealis AK	DHK0334b	USA, Alaska, Kodiak Island, Buskin River, 20 m, 57.76541°N, 152.52016°W	CAS
Pelophila borealis RFE	DHK2064	Russia, Kamtschatka, os. Asabatsch'e and river Kamtschatka, 56.22278°N, 162.01917°E	CAS
Pelophila rudis	DRM2376	USA, Alaska, Fairbanks, east shore of Smith Lake, 150 m, 64.8622°N, 147.8616°W	OSAC
outgroup specimens
Bembidion antiquum	DRM1963	USA, Missouri, Carter County, Current River at Van Buren, 135 m, 36.99037°N, 91.01672°W	OSAC
Calosoma scrutator	DHK1712	USA, Oklahoma, Sequoyah County, Sallisaw, 245 m, 35.53371°N, 94.62467°W	CAS
Calosoma scrutator	DRM2249	USA, Arizona, Santa Cruz County, Pena Blanca, 31.3852°N, 111.0931°W	OSAC
Loricera pilicornis	DHK1185	Canada, Newfoundland, Deer Lake, north shore, 5 m, 49.18058°N, 57.45150°W	CAS
Metrius contractus	DHK1424	USA, California, Mendocino County, Angelo Reserve, Skunk Creek, 420 m, 39.72554°N, 123.64412°W	CAS
Pterostichus melanarius	DRM2311	USA, Wisconsin, Dane County, Madison, 43.086°N, 89.425°W	OSAC
Scaphinotus petersi	DRM0878	USA, Arizona, Pima County, Mt. Lemmon Ski Valley, 2500 m, 32.447°N, 110.777°W	OSAC
Trachypachus inermis	DHK0738a	USA, California, Tuolumne County, Sonora Pass, 2920 m, 38.32961°N, 119.63857°W	CAS
Trachypachus slevini	DHK0513a	USA, Oregon, Lincoln County, Moolack Beach, 8 m, 44.71078°N, 124.06035°W	CAS