Systematic assessment of the Leporinus desmotes species complex, with a description of two new species Author Burns, Michael D. Oregon State University, Department of Fisheries and Wildlife, 104 Nash Hall Corvallis, OR 97330 michael.burns@oregonstate.edu Author Chatfield, Marcus Oregon State University, Department of Fisheries and Wildlife, 104 Nash Hall Corvallis, OR 97330 marcus.chatfield@oregonstate.edu Author Birindelli, José L. O. Departamento de Biologia Animal e Vegetal, Universidade Estadual de Londrina, Londrina, PR, Brazil josebirindelli@yahoo.com Author Sidlauskas, Brian L. Oregon State University, Department of Fisheries and Wildlife, 104 Nash Hall Corvallis, OR 97330 brian.sidlauskas@oregonstate.edu text Neotropical Ichthyology 2017 2017-07-03 15 2 e 160166 1 24 http://dx.doi.org/10.1590/1982-0224-20160166 journal article 315423 10.1590/1982-0224-20160166 a556348c-05b1-49db-afe5-5c7ad8ad0604 1982-0224 20632953

Tab. 6. Linear morphometrics for all examined species summarized as observed ranges, with mean ± standard deviation in parentheses. Measures 1 to 29 are percentages of SL, measures 30-37 are percentages of head length.

		L. enyae (holotype)	L. enyae (paratypes)	L. villasboasorum (holotype)	L. villasboasorum (paratypes)	L. desmotes	L. jatuncochi
	Standard length (mm)	145.6	37.7 - 181.4 (110.4 ± 39.9)	91.0	69.9 - 127.9 (97.7 ± 19.6)	56.4 - 170.8 (96.2 ± 46.5)	22.8 - 84.8 (58.5 ± 19.3)
1	Snout to dorsal-fin origin	43.4	41.9 - 47.2 (44.3 ± 1.3)	46.7	43.6 - 46.7 (45.3 ± 1.1)	42.6 - 46.0 (44.5 ± 1.1)	41.8 - 49.2 (44.8 ± 1.9)
2	Snout to adipose-fin origin	82.0	78.7 - 83.7 (81.4 ± 1.3)	80.9	80.1 - 82.8 (81.0 ± 0.9)	80.1 - 82.9 (81.5 ± 0.9)	78.6 - 83.0 (81.0 ± 1.1)
3	Snout to anal-fin origin	78.8	75.4 - 79.7 (77.9 ± 1.2)	76.9	74.6 - 78.0 (76.3 ± 1.2)	76.8 - 79.5 (77.9 ± 0.9)	75.1 - 80.5 (77.0 ± 1.4)
4	Snout to pelvic-fin origin	46.2	45.4 - 49.5 (47.1 ± 1.0)	47.8	45.9 - 50.0 (48.4 ± 1.4)	45.8 - 50.1 (48.4 ± 1.5)	44.8 - 52.1 (47.5 ± 2.4)
5	Snout to pectoral-fin origin	24.3.	22.3 - 28.2 (25.2 ± 1.2)	27.2	24.7 - 28.6 (27.3 ± 1.3)	24.0 - 27.3 (25.8 ± 1.4)	25.1 - 33.4 (27.6 ± 2.2)
6	Dorsal-fin origin to pectoral-fin origin	27.9	25.4 - 30.5 (27.8 ± 1.2)	28.2	26.1 - 28.8 (27.4 ± 0.9)	27.3 - 30.9 (29.0 ± 1.3)	21.8 - 30.3 (27.6 ± 1.9)
7	Dorsal-fin origin to pelvic-fin origin	26.4	22.1 - 28.3 (25.6 ± 1.4)	25.3	22.7 - 25.3 (24.4 ± 0.9)	23.1 - 29.7 (26.7 ± 2.1)	22.2 - 30.1 (27.1 ± 1.9)
8	Dorsal-fin origin to anal-fin origin	43.8	40.1 - 43.8 (42.1 ± 1.0)	40.6	39.1 - 41.3 (40.3 ± 0.9)	40.2 - 46.2 (43.1 ± 1.9)	38.3 - 44.1 (42.2 ± 1.5)
9	Dorsal-fin origin to anal-fin insertion	49.6	33.9 - 54.4 (47.6 ± 3.2)	46.9	44.5 - 48.7 (46.3 ± 1.3)	46.5 - 52.7 (49.7 ± 2.0)	43.9 - 51.0 (48.3 ± 2.1)
10	Dorsal-fin origin to posterior margins of the hypurals	61.1	56.0 - 62.4 (59.5 ± 1.4)	58.7	56.5 - 60.5 (59.0 ± 1.3)	58.0 - 62.8 (60.3 ± 1.5)	55.3 - 62.2 (59.7 ± 1.7)
11	Dorsal-fin origin to adipose-fin origin	41.4	36.2 - 42.6 (39.8 ± 1.5)	38.0	36.7 - 39.4 (38.4 ± 0.9)	36.6 - 43.1 (40.3 ± 1.9)	35.7 - 42.1 (39.7 ± 1.6)
12	Length of dorsal-fin base	13.7	11.7 - 15.5 (13.9 ± 0.8)	15.5	14.4 - 15.6 (15.0 ± 0.4)	15.4 - 16.9 (16.0 ± 0.5)	13.8 - 17.0 (15.6 ± 0.9)
13	Dorsal-fin insertion to pelvic-fin origin	24.0	20.9 - 26.7 (23.8 ± 1.3)	25.0	22.1 - 25.0 (23.1 ± 0.9)	22.3 - 28.2 (24.9 ± 1.7)	20.4 - 27.9 (24.8 ± 2.1)
14	Dorsal-fin insertion to adipose-fin origin	28.1	22.9 - 28.7 (26.1 ± 1.5)	20.8	20.8 - 26.5 (23.7 ± 1.6)	21.7 - 26.6 (24.1 ± 1.5)	20.2 - 26.9 (24.2 ± 1.7)
15	Dorsal-fin insertion to anal-fin origin	30.9	27.3 - 31.3 (29.3 ± 1.1)	27.7	25.9 - 29.3 (27.1 ± 1.1)	25.2 - 31.5 (28.4 ± 1.9)	24.5 - 30.1 (27.9 ± 1.8)
16	Dorsal-fin insertion to anal-fin insertion	35.7	31.7 - 36.4 (34.3 ± 1.2)	31.4	30.4 - 33.4 (31.6 ± 1.0)	30.0 - 35.5 (33.6 ± 1.8)	29.3 - 36.0 (33.2 ± 1.8)
17	Adipose-fin origin to anal-fin origin	17.3	15.1 - 19.1 (16.9 ± 1.0)	17.1	16.3 - 17.4 (17.0 ± 0.4)	17.9 - 19.8 (18.7 ± 0.7)	15.9 - 20.8 (18.3 ± 1.2)
18	Adipose-fin origin to anal-fin insertion	13.0	12.1 - 14.8 (13.1 ± 0.7)	12.5	12.5 - 13.6 (13.1 ± 0.4)	13.9 - 15.6 (14.7 ± 0.6)	11.6 - 15.6 (14.2 ± 1.1)
19	Adipose-fin origin to posterior margins of the hypurals	19.9	18.3 - 22.1 (19.7 ± 0.8)	20.8	19.7 - 21.8 (20.6 ± 0.6)	18.6 - 20.9 (20.2 ± 0.7)	18.3 - 21.8 (20.0 ± 1.0)
20	Length of anal-fin base	9.5	8.0 - 11.3 (9.4 ± 0.9)	9.3	9.0 - 10.0 (9.3 ± 0.4)	9.0 - 12.1 (10.5 ± 1.1)	10.0 - 13.2 (11.6 ± 1.0)
21	Anal-fin insertion to posterior margin of hypurals	16.3	13.9 - 17.2 (15.4 ± 0.8)	16.7	15.1 - 18.4 (16.7 ± 1.1)	13.9 - 16.1 (14.8 ± 0.6)	12.2 - 16.2 (14.8 ± 1.1)
22	Pelvic-fin origin to anal-fin origin	33.4	29.0 - 36.3 (32.2 ± 1.6)	29.5	27.5 - 31.4 (29.2 ± 1.2)	28.7 - 34.4 (30.7 ± 2.1)	24.8 - 34.5 (30.6 ± 2.5)
23	Pelvic-fin origin to adipose-fin origin	42.0	36.7 - 43.7 (40.4 ± 1.5)	38.4	36.3 - 38.6 (37.7 ± 0.9)	38.5 - 43.4 (40.2 ± 1.7)	34.8 - 43.2 (40.1 ± 1.9)
24	Pelvic origin to posterior margin of hypurals	58.5	53.2 - 58.5 (55.9 ± 1.2)	55.3	53.7 - 55.7 (54.6 ± 0.8)	52.9 - 57.1 (55.0 ± 1.6)	50.8 - 57.5 (55.3 ± 2.0)
25	Pelvic-fin origin to pectoral-fin origin	22.5	20.4 - 24.9 (22.4 ± 1.1)	22.7	20.9 - 23.4 (22.5 ± 0.9)	21.5 - 24.9 (23.1 ± 1.1)	18.4 - 26.7 (21.1 ± 1.8)
26	Greatest body depth	26.0	23.2 - 28.7 (25.4 ± 1.4)	24.4	23.0 - 24.9 (24.0 ± 0.6)	25.1 - 28.9 (26.7 ± 1.3)	24.2 - 29.7 (27.0 ± 1.6)
27	Greatest body width	12.2	10.3 - 13.9 (11.9 ± 1.0)	12.6	11.2 - 13.2 (12.6 ± 0.7)	11.3 - 15.5 (13.2 ± 1.5)	10.8 - 14.2 (12.4 ± 1.0)
28	Caudal-peduncle depth	9.4	8.1 - 10.9 (9.5 ± 0.6)	9.7	9.1 - 10.3 (10.0 ± 0.4)	10.1 - 11.2 (10.6 ± 3.7)	8.6 - 11.6 (10.4 ± 0.8)
29	Head length	21.6	21.6 - 26.3 (23.4 ± 1.0)	24.8	24.1 - 26.0 (25.1 ± 0.7)	22.5 - 26.3 (24.9 ± 1.5)	23.9 - 30.1 (26.4 ± 1.9)
30	Preopercle length	82.1	76.1 - 83.7 (80.8 +- 1.8)	79.7	78.8 - 82.6 (80.0 +- 1.2)	77.9 - 82.0 (79.8 ± 1.2)	72.1 - 82.9 (78.2 ± 2.7)
31	Snout to anterior margin of the eye	43.0	37.2 - 43.9 (41.4 +- 1.5)	38.7	38.7 - 43.6 (40.8 +- 1.9)	36.8 - 42.4 (40.8 ± 1.9)	32.6 - 45.7 (39.1 ± 3.4)
32	Head depth	78.8	69.9 - 78.9 (73.9 +- 2.7)	73.6	71.2 - 80.9 (75.1 +- 3.1)	67.8 - 78.7 (72.8 ± 3.6)	60.8 - 78.4 (70.9 ± 4.5)
33	Snout depth	53.2	44.5 - 53.2 (47.9 +- 2.2)	49.1	47.8 - 56.5 (50.0 +- 3.0)	42.9 - 51.7 (47.5 ± 3.1)	42.3 - 49.7 (45.5 ± 2.4)
34	Jaw length	24.1	20.1 - 26.8 (23.5 +- 1.9)	22.2	21.1 - 25.1 (23.0 +- 1.4)	20.7 - 27.2 (23.4 ± 2.7)	18.4 - 26.9 (21.5 ± 2.2)
35	Eye diameter	27.4	23.7 - 32.9 (28.1 +- 2.7)	28.8	22.7 - 29.7 (26.9 +- 2.1)	25.1 - 34.9 (29.5 ± 3.1)	27.1 - 37.0 (31.4 ± 3.1)
36	Interorbital width	43.3	35.2 - 44.7 (40.6 +- 2.2)	38.9	35.8 - 41.7 (38.7 +- 1.8)	40.9 - 47.8 (43.9 ± 2.5)	35.8 - 44.1 (39.7 ± 2.5)
37	Snout to supraoccipital crest	92.2	82.9 - 1.0 (87.6 +- 4.0)	93.1	80.9 - 93.1 (88.1 +- 4.0)	79.4 - 86.7 (83.5 ± 2.6)	76.3 - 89.4 (82.8 ± 3.5)