S. campiaoae sp. nov. S. campiaoae sp. nov. S. dixoni S. dixoni S. heliocostai S. heliocostai S. scelopori S. scelopori S. spinosulus S. spinosulus Male Female Male Female Male Female Male Female Male Female Length 1036 ± 143 (896– 3007 ± 966 (677– 2320 ± 340 (1920– 6670 ± 770 (5440– 1020– 2190– 1960 3810–5730 1470– 7440– 1161) 3668) 2800) 7870) 1260 8400 1720 8960 Width 101 ± 25 (75–134) 144 ± 29 (76–167) 279 ± 55 (204–357) 751 ± 100 (638–969) 140–150 140–160 136 3810–5980 140–150 140–150 Esophagus total 210 ± 30 (165–227) 479 ± 52 (373–531) 210 ± 30 (165–227) 479 ± 52 (373–531) - - - - - - - - Esophagus length 155 ± 26 (116–174) 370 ± 32 (306–414) 155 ± 26 (116–174) 370 ± 32 (306–414) 221 ± 24 (180–250) 445 ± 47 (372–512) 190–210 300–510 285 4900–5170 180–200 410–430 Isthmus 9 10 25 ± 6 (18–34) 10 ± 2 (6–12) Bulb length 57 ± 7 (46–63) 103 ± 20 (64–124) 80 ± 9 (67–92) 131 ± 17 (110–159) 54–57 57–90 72 1050–1080 39–60 75–100 Bulb width 63 ± 11 (47–70) 114 ± 22 (74–142) 79 ± 10 (61–88) 135 ± 25 (92–171) 46–61 55–100 72 1200–1290 50–72 90–130 Nerve ring 60 ± 5 (54–64) 107 ± 28 (63–130) 123 ± 16 (92–134) 123 ± 15 (92–140) 79–90 93–150 132 1090–1200 43–50 100–130 Excretory pore 306 168 790 ± 122 (638–995) 828 ± 86 (740–1020) 168 790 ± 122 (638–995) 828 ± 86 (740–1020) 168 790 ± 122 (638–995) 828 ± 86 (740–1020) 280 190–280 449 5710–6530 300–560 Tail length 216 ± 70 (117–275) 577 ± 136 (291– 216 ± 70 (117–275) 577 ± 136 (291– 380 ± 23 (357–408) 258 ± 32 (204–306) 190 140–390 81 8570– 140–180 450–630 731) 11.560 Spicule - - - - - - 57 - 43–50 - Distance of vulva from - Prebulbar - Postbulbar - Prebulbar - Postbulbar - Postbulbar anterior end 158 ± 46 915 ± 106 (816– 200– 6250– 370– (61–206) 1148) 300 7210 600 Caudal spines 3–9 18–51 1–3 3 - - - 12–14 - 70–80 Anus - 231 ± 152 (67–366) - 231 ± 152 (67–366) - - - 630–770 - - - 1750– 1890 Egg length - 114 ± 6 (102–124) - 91 ± 3 (88–95) 140 1620–1710 160–180 Egg width - 37 ± 8 (19–49) - 41 ± 1 (40–43) - 54 - 36–48 - 36–50 Caudal papillae 4 pairs - 3 pairs - 3 pairs - 4 - 3 pairs - pairs