An integrative redescription of Echiniscus quadrispinosus quadrispinosus Richters, 1902 (Heterotardigrada, Echiniscidae) from the terra typica in Taunus Mountain Range (Europe; Germany) Author Kaczmarek, Łukasz FE20F8A0-B5E8-4C17-869C-29AA12DFF672 Department of Animal Taxonomy and Ecology, Faculty of Biology, Adam Mickiewicz University in Poznań, Uniwersytetu Poznańskiego 6, 61 - 614 Poznań, Poland. kaczmar@amu.edu.pl Author Kayastha, Pushpalata C09EF655-F729-4A3E-90BF-D441F0B2F957 Department of Animal Taxonomy and Ecology, Faculty of Biology, Adam Mickiewicz University in Poznań, Uniwersytetu Poznańskiego 6, 61 - 614 Poznań, Poland. kaypus@amu.edu.pl Author Gawlak, Magdalena E1252646-58B9-4AF4-89EA-2E602D979FD2 The Institute of Plant Protection-National Research Institute, Węgorka 20, 60 - 318 Poznań, Poland. m.gawlak@iorpib.poznan.pl Author Mioduchowska, Monika 0699908B-B303-4220-A91D-BB1431158EF1 Department of Evolutionary Genetics and Biosystematics, Faculty of Biology, University of Gdansk, Wita Stwosza 59, 80 - 308 Gdansk, Poland. & Department of Marine Plankton Research, Institute of Oceanography, University of Gdansk, Marszałka Piłsudskiego 46, 81 - 378 Gdynia, Poland. & Department of Invertebrate Zoology and Hydrobiology, Faculty of Biology and Environmental Protection, University of Lodz, Banacha 12 / 16, 90 - 237 Lodz, Poland. monika.mioduchowska@ug.edu.pl Author Roszkowska, Milena 3C2055BD-866C-4B74-8669-A0749C6C9D7D Department of Animal Taxonomy and Ecology, Faculty of Biology, Adam Mickiewicz University in Poznań, Uniwersytetu Poznańskiego 6, 61 - 614 Poznań, Poland. & Department of Bioenergetics, Faculty of Biology, Adam Mickiewicz University in Poznań, Uniwersytetu Poznańskiego 6, 61 - 614, Poznań, Poland. roszkowska@gmail.com text European Journal of Taxonomy 2022 2022-06-14 823 1 102 124 http://zoobank.org/8824e8c0-ee82-4803-8480-e4fd2773fc80 journal article 77866 10.5852/ejt.2022.823.1819 23307e3f-3bd6-47e7-8d8e-21e272ba6691 2118-9773 6658659 8824E8C0-EE82-4803-8480-E4FD2773FC80

Table 1. Measurements (in µm) and sp values of selected morphological structures of adult females from the neotype population of Echiniscus q. quadrispinosus Richters, 1902 mounted in Hoyer’s medium. Abbreviations: N = number of specimens/structures measured; RANGE = refers to the smallest and the largest structure among all measured specimens; SD = standard deviation.

Character	N	Range		Mean		SD		Neotype		N	Range		Mean		SD
		µm	sp	µm	sp	µm	sp	µm	sp		µm	sp	µm	sp	µm	sp
Body length	20	204–280	450–562	241	502	25	24	245	507	20	190–236	469 –536	219	504	12	15
Scapular plate length	20	40.0–60.2	–	48.0	–	5.3	–	48.3	–	20	39.3–46.2	–	43.4	–	1.9	–
Head appendage lengths										0
Cirrus internus	19	18.5–26.8	42.2–55.2	22.3	46.8	2.1	4.5	21.4	44.4	20	15.8–20.4	37.7 –45.0	18.2	41.9	1.5	2.4
Cephalic papilla	20	6.4–10.3	13.9–23.3	8.7	18.1	1.2	2.1	9.6	19.9	20	6.7–8.8	15.7 –20.6	7.6	17.4	0.6	1.5
Cirrus externus	20	23.2–32.0	51.2–64.6	26.9	56.0	3.1	4.1	29.6	61.2	19	18.4–23.5	43.9 –51.9	20.8	47.8	1.5	2.7
Clava	20	5.6–7.4	12.0–16.4	6.6	13.9	0.5	1.3	7.0	14.4	20	4.4–6.8	10.1 –16.2	5.7	13.3	0.6	1.5
Cirrus A	19	46.3–56.9	87.6–121.4	52.4	109.6	3.1	9.8	56.9	117.8	20	40.2–52.7	92.8 –120.6	46.6	107.3	3.9	9.0
Cirrus A /Body length ratio	19	19%–25%	–	22%	–	2%	–	23%	–	20	18%–24%	–	21%	–	2%	–
Cirrus int/ext length ratio	19	71%–92%	–	84%	–	6%	–	72%	–	19	81%–95%	–	88%	–	4%	–
Body appendage lengths										0
Cirrus B	18	35.8–77.9	70.3–160.6	57.1	118.5	10.7	17.9	57.3	118.7	19	19.3–43.7	45.3 –97.9	33.6	77.5	6.5	15.5
Cirrus C	20	57.8–92.1	125.5–181.9	70.5	147.1	9.9	15.5	72.8	150.6	20	42.7–67.4	117.4 –151.4	56.6	130.4	4.5	9.3
Cirrus Cd	20	17.1–30.6	40.0–63.0	24.2	50.6	2.6	5.5	22.4	46.4	20	16.9–26.6	39.5 –54.2	20.8	48.0	2.0	4.5
Cirrus D	15	45.4–73.0	93.9–137.5	55.7	118.5	8.4	12.2	54.4	112.7	18	32.8–58.3	83.7 –110.7	42.9	98.9	5.0	9.3
Cirrus Dd	19	10.8–22.8	25.9–47.0	17.0	35.4	2.7	4.9	17.3	35.7	18	13.5–20.5	30.3 –43.2	15.9	36.5	1.8	3.8
Cirrus E	18	46.7–114.4 109.4–190.0		77.1	159.0	13.8	16.0	73.3	151.8	18	44.0–68.0	118.9 –158.8	60.9	139.7	5.0	12.6
Spine on leg I length	19	3.4–5.4	6.2–11.1	4.3	8.9	0.6	1.2	4.3	8.9	16	3.1–4.6	6.9 –10.5	3.8	8.6	0.5	1.1
Papilla on leg IV length	18	3.7–5.2	7.7–10.7	4.5	9.4	0.5	1.0	3.7	7.7	18	3.3–4.9	8.2 –10.7	4.0	9.2	0.4	0.8
Number of teeth on the collar	19	7–10	–	8.1	–	1.0	–	8	–	18	7–10	–	8.4	–	1.0	–
Notch length	19	12.2–18.1	28.1–34.6	15.4	32.4	1.7	1.7	15.6	32.3	16	12.0–15.6	27.0 –36.0	13.5	30.8	1.1	2.4
Claw 1 lengths										0
Claw	19	12.0–18.4	27.7–36.2	14.9	31.3	1.7	2.6	16.0	33.0	19	11.5–14.0	26.2 –33.3	12.6	29.0	0.7	2.0
Spur	15	1.9–4.4	4.2–9.1	2.9	6.1	0.7	1.7	4.4	9.1	12	1.3–2.2	3.2 –4.8	1.8	4.2	0.3	0.5
Spur/claw length ratio	15	15%–28%	–	20%	–	5%	–	0	–	12	11%–17%	–	15%	–	2%	–
Claw 2 lengths										0
Claw	17	10.2–16.9	24.5–32.0	13.8	28.8	1.9	2.2	14.2	29.4	17	10.0–13.2	23.7 –29.8	12.0	27.5	0.6	1.6
Spur	11	2.1–3.0	4.0–6.6	2.5	5.2	0.2	0.9	?	?	9	1.5–2.3	4.0 –5.4	2.0	4.5	0.2	0.4
Spur/claw length ratio	11	14%–25%	–	19%	–	4%	–	?	–	9	14%–20%	–	17%	–	2%	–
Claw 3 lengths										0
Claw	20	11.1–17.5	25.4–36.1	14.1	29.5	2.0	3.0	14.2	29.4	17	11.0–13.4	25.1 –29.3	12.1	27.6	0.4	1.1
Spur	15	1.8–3.1	3.6–6.2	2.4	5.0	0.3	0.7	?	?	9	1.7–2.1	3.9 –4.6	1.9	4.4	0.1	0.3
Spur/claw length ratio	15	14%–21%	–	17%	–	3%	–	?	–	9	13%–18%	–	16%	–	1%	–
Claw 4 lengths										0
Claw	20	13.3–19.3	30.8–38.4	16.4	34.2	1.6	2.2	16.4	34.0	19	12.6–15.7	30.2 –36.4	14.3	32.9	0.8	1.8
Spur	13	2.1–4.2	4.8–7.8	2.8	5.8	0.6	1.1	?	?	5	1.6–2.6	4.3 –5.8	2.2	5.0	0.3	0.6
Spur/claw length ratio	13	15%–24%	–	17%	–	4%	–	?	–	5	13%–18%	–	16%	–	2%	–