Revision of the " Chloritis delibrata (Benson, 1836) " group (Gastropoda, Stylommatophora, Camaenidae) Author Pall-Gergely, Barna https://orcid.org/0000-0002-6167-7221 Plant Protection Institute, Centre for Agricultural Research, Herman Otto Street 15, Budapest, H- 1022, Hungary pallgergely2@gmail.com Author Ablett, Jonathan D. https://orcid.org/0000-0002-7277-1934 Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, UK Author Szabo, Marton Hungarian Natural History Museum, Department of Paleontology and Geology, Ludovika ter 2, Budapest 1083, Hungary Author Neubert, Eike https://orcid.org/0000-0002-0277-2894 Natural History Museum of the Burgergemeinde Bern, Bernastr. 15, CH- 3005 Berne, Switzerland & Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland text ZooKeys 2022 2022-02-15 1086 1 31 http://dx.doi.org/10.3897/zookeys.1086.77180 journal article http://dx.doi.org/10.3897/zookeys.1086.77180 1313-2970-1086-1 0AFF7CFB9400477F81160120C393C4FD BD013F15E39C5B5FBE666837836810E1
Table 1. Most important character states of camaenid genera relevant for this study.
ReferencesPenisInner structure of the penisPenial sheathEpiphallusInsertion of retractor musclePenial caecumFlagellumAdditional organ
Bouchetcamaena Thach, 2018this studylong, apically thickenedparallel folds and a vestigial vergeabsentlong, cylindricalon distal epiphallusabsentlong, slenderabsent
Continental " Chloritis "Sutcharit and Panha (2010), Pall-Gergely et al. (2020)long, apically thickenedparallel folds and a large vergeabsentlong, cylindricalon distal epiphallusabsentmedium length, gradually becoming slenderabsent
Neotrachia Schileyko, 2018Schileyko (2018)short, thicklongitudinal pilasters, broken into series of tuberclesabsentswollen, ovoidpenis-epiphallus transitionabsentmedium length, gradually becoming slenderabsent
Trachia Martens, 1860 (based on T. vittata)Schileyko (2003)short, swollenchaotically arranged pilasterscovers entire penisrather short, thickincorporated into penial sheathabsentrather long, gradually becoming slender-
Trichochloritis Pilsbry, 1891Collinge (1903), Pall-Gergely and Neubert (2019)long, apically thickenedunknownabsentlong, cylindricalon distal epiphallusmoderately long, slendervery short, pointedabsent
Burmochloritis Godwin-Austen, 1920Godwin-Austen (1920), unpublished informationlong, thick, cylindricalwavy folds, verge absentabsentlong, cylindricalbounds penis and epiphallus at some distance from their junctionshort, pointedlong, slenderlong, cylindrical, derives from wall of vagina
Satsuma A. Adams, 1868Wang et al. (2014), Zhang et al. (2020)long, cylindricalwavy folds, verge absentabsentlong, cylindricalon distal epiphalluswell-developed, taperinglong to shortabsent
Sinochloritis M. Wu & Z. Chen, 2019Wu & Chen, 2019thick, cylindricalparallel folds, verge absentabsentlong, cylindricalon distal epiphallus, and also covers proximal part of penislarge, internally with "peach shaped epiphallic papilla"long, slender, taperingabsent
Bellatrachia Schileyko, 2018Schileyko (2018), Pall-Gergely and Neubert (2019)long, cylindricalparallel foldsabsentlong, cylindricalpenis-epiphallus transitionabsentthick, somewhat swollen, with slender tipabsent
Planispira Beck, 1837Schileyko (2003)short, apically thickenedfolds and an ovoid, large vergeabsentlong, cylindricalmiddle of epiphallusabsentshort, conicalabsent