Phylogeographic pattern and taxonomic revision of the Kaloula baleata species complex (Amphibia, Anura, Microhylidae) with description of two new species from Indochina Author Gorin, Vladislav A. 0000-0002-9017-1502 Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, Moscow 119234, Russia Author Orlov, Nikolai L. 0000-0003-4401-348X Division of Herpetology & Ormithology, Zoological Institute, Russian Academy of Sciences, St. Petersburg 199034 Universitetskaya nab., 1, 33701, Russia Author Bragin, Andrey M. 0000-0002-3621-9763 Joint Vietnam - Russia Tropical Science & Technology Research Center, 63 Nguyen Van Huyen Road, Nghia Do, Cau Giay, Hanoi 122000, Vietnam Author Pawangkhanant, Parinya 0000-0002-0947-5729 Division of Fishery, School of Agriculture & Natural Resources, University of Phayao, Phayao 56000, Thailand & Rabbit in the Moon Foundation, Suanphueng, Ratchaburi 70180, Thailand Author Milto, Konstantin D. 0000-0003-4061-0164 Division of Herpetology & Ormithology, Zoological Institute, Russian Academy of Sciences, St. Petersburg 199034 Universitetskaya nab., 1, 33701, Russia Author Le, Dac Xuan 0000-0003-4586-6214 Joint Vietnam - Russia Tropical Science & Technology Research Center, 63 Nguyen Van Huyen Road, Nghia Do, Cau Giay, Hanoi 122000, Vietnam Author Nguyen, Tan Van 0000-0001-5413-968X Institute for Research & Training in Medicine, Biology & Pharmacy, Duy Tan University, Da Nang 550000, Vietnam Author Dufresnes, Christophe 0000-0002-8497-8908 College of Medicine & Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang 550000, Vietnam Author Suwannapoom, Chatmongkon 0000-0002-3342-1464 Division of Fishery, School of Agriculture & Natural Resources, University of Phayao, Phayao 56000, Thailand Author Poyarkov, Nikolay A. 0000-0002-7576-2283 Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, Moscow 119234, Russia & Joint Vietnam - Russia Tropical Science & Technology Research Center, 63 Nguyen Van Huyen Road, Nghia Do, Cau Giay, Hanoi 122000, Vietnam text Herpetozoa 2024 2024-12-11 37 391 420 journal article 306388 10.3897/herpetozoa.37.e137394 5be396ed-af3c-4af5-b187-1c48444979fb E97A128A-13B5-4781-8FEC-ADC5CB37AB45

Table 4. Measurements (in mm) of specimens of Kaloula discordia sp. nov., Kaloula laosensis sp. nov., and K. indochinensis. For character abbreviations, see the Materials and methods section. Abbreviations: F – adult female; M – adult male; SF – subadult female; SM – subadult male.

Species	Sample ID	Sex	Status	SVL	HL	SL	EL	N-EL	HW	IND	IOD	UEW	FLL	LAL	HAL	1 FL	IPTL	OPTL	3 FDD	HLL	TL	FL	OMTL	1 TOEL	4 TDD
Kaloula discordia sp. nov.	ZMMU - A- 8134	M	holotype	51.5	11.7	5.5	4.1	3.9	14.8	3.2	5.3	3.0	31.9	26.4	15.7	8.1	3.0	3.5	3.6	60.0	14.7	30.2	2.1	7.9	2.3
K. discordia sp. nov.	ZMMU - A- 4739	M	paratype	52.0	11.9	5.2	4.0	3.9	15.2	3.2	5.3	2.9	33.4	26.8	15.8	8.2	3.0	3.4	3.8	62.7	15.3	31.4	2.5	7.7	2.9
K. discordia sp. nov.	ZISP 15285	M	paratype	48.0	10.8	5.1	3.1	3.7	13.7	3.1	5.0	2.8	31.2	24.1	15.0	7.8	2.9	3.3	3.2	58.0	13.4	28.9	2.5	7.9	2.6
K. discordia sp. nov.	ZMMU - A- 8135	M	paratype	49.6	11.3	5.3	3.4	3.8	14.0	3.2	4.9	2.7	31.5	25.3	15.3	7.9	2.6	3.3	3.5	59.0	14.1	29.7	2.4	7.6	2.8
K. discordia sp. nov.	ZMMU - A- 8136	M	paratype	45.4	10.6	4.9	3.3	3.4	13.8	3.0	4.8	2.7	31.4	24.2	14.6	7.8	2.9	2.8	3.3	57.4	14.1	27.9	2.2	7.2	2.4
K. discordia sp. nov.	ZMMU - A- 8137	M	paratype	42.9	10.3	4.9	3.0	3.0	13.7	3.0	4.5	2.6	30.8	24.0	14.9	7.8	2.3	3.1	3.0	56.3	13.1	28.5	2.0	7.1	2.4
K. discordia sp. nov.	ZMMU - A- 8138	M	paratype	48.7	11.1	5.3	3.7	3.5	14.7	3.2	4.9	2.9	31.1	25.0	15.4	7.9	2.4	3.0	3.5	60.6	14.4	30.5	2.2	7.4	2.3
K. discordia sp. nov.	ZMMU - A- 8139	M	paratype	49.5	11.2	5.1	3.1	3.5	15.0	3.1	4.7	3.0	31.7	24.9	14.2	7.4	2.6	3.1	3.3	60.5	13.8	31.0	1.9	7.5	2.7
K. discordia sp. nov.	ZMMU - A- 8140	M	paratype	47.2	10.5	5.0	3.4	3.3	15.1	3.0	4.9	2.7	27.9	23.0	14.1	6.8	2.7	3.3	3.5	57.6	14.7	27.4	1.9	6.9	2.8
K. discordia sp. nov.	ZISP 15286	M	paratype	52.6	12.2	5.8	3.8	3.8	15.7	3.2	5.6	2.8	32.9	26.3	15.6	7.9	3.2	3.6	4.0	61.9	15.1	31.1	2.3	8.2	3.4
K. discordia sp. nov.	ZISP 15287	M	paratype	49.1	10.6	5.3	3.6	3.5	14.5	3.1	5.0	2.7	31.2	24.6	15.4	8.1	2.8	3.1	3.4	60.6	14.3	29.7	2.6	8.2	2.6
K. discordia sp. nov.	ZMMU - A- 8141	M	paratype	50.0	11.3	5.5	3.5	3.7	15.3	3.2	5.0	3.0	31.8	25.2	15.2	7.5	2.6	3.0	3.3	60.5	14.3	31.0	1.9	7.4	2.6
K. discordia sp. nov.	ZMMU - A- 8142	M	paratype	52.2	11.7	5.1	3.6	3.2	15.3	3.2	5.0	2.8	31.9	25.1	15.6	7.9	2.5	3.0	3.6	61.0	14.1	30.8	1.9	7.2	2.7
K. discordia sp. nov.	ZMMU - A- 8143	M	paratype	51.5	11.7	5.5	3.4	3.6	15.0	3.2	5.1	2.7	32.1	25.4	15.2	7.9	2.9	3.1	3.7	58.5	13.4	29.9	2.4	7.5	2.6
K. discordia sp. nov.	ZMMU - A- 4642	F	paratype	56.2	12.8	6.2	4.1	4.2	17.0	3.8	5.9	3.0	35.9	28.2	16.3	9.3	3.2	3.7	4.0	67.4	16.7	33.3	3.0	9.0	3.0
K. discordia sp. nov.	ZMMU - A- 4602-1	F		56.4	13.0	5.4	4.2	4.0	16.9	3.7	6.0	3.0	35.8	28.6	16.6	10.0	3.4	3.4	3.8	67.6	17.2	33.7	3.0	9.1	2.9
K. discordia sp. nov.	ZMMU - A- 4602-2	SF		25.3	7.0	3.1	2.5	2.8	7.5	1.9	3.2	1.5	15.5	11.9	7.0	3.8	2.0	1.8	1.7	30.2	7.4	13.9	-	3.5	1.5
K. discordia sp. nov.	ZMMU - A- 4602-3	SF		35.0	8.3	3.8	3.0	2.8	10.2	2.6	3.6	1.9	23.2	14.5	10.3	5.9	2.2	2.2	2.6	40.2	9.6	20.4	1.4	4.5	2.0
Kaloula laosensis sp. nov.	ZISP 15284	M	holotype	44.0	10.6	4.6	3.3	3.2	12.6	2.7	4.3	2.4	27.6	21.7	13.0	7.5	2.7	2.8	3.4	49.3	11.6	25.4	2.0	7.3	2.2
K. laosensis sp. nov.	ZMMU - A- 8144	F	paratype	54.8	12.4	5.3	5.0	5.2	16.5	3.6	5.7	3.0	33.8	28.3	17.8	10.4	3.2	3.7	4.3	66.9	16.4	32.5	2.8	8.4	2.8
Kaloula indochinensis	ZMMU - A- 8147	M		45.3	11.0	4.9	3.9	3.3	14.7	3.1	5.0	2.8	31.5	23.7	13.9	8.1	3.1	3.4	3.8	55.2	14.0	27.2	2.19	7.2	2.3
K. indochinensis	ZMMU - A- 8148	M		50.9	11.6	5.3	3.4	3.8	15.1	3.0	4.5	2.5	32.5	24.8	14.6	7.9	2.9	3.1	3.3	57.3	14.5	27.5	2.72	8.1	2.4
K. indochinensis	ZMMU - A- 8149	M		42.3	9.8	4.8	3.4	3.3	12.0	2.9	4.5	2.4	28.1	20.4	12.6	7.8	2.4	2.6	2.9	47.9	10.9	24.9	2.15	6.2	2.2
K. indochinensis	ZMMU - A- 8150	M		46.3	10.7	4.9	3.5	3.5	13.0	3.1	4.5	2.6	29.9	22.4	13.9	7.9	2.6	3.0	3.4	50.7	12.2	26.2	2.59	7.0	2.4
K. indochinensis	ZMMU - A- 8151	M		48.5	11.2	5.2	3.9	3.5	15.7	3.1	4.7	2.7	31.1	24.0	14.6	8.2	3.2	3.3	3.7	55.1	12.9	27.6	2.44	7.4	2.7
K. indochinensis	ZMMU - A- 8152	M		49.1	11.9	5.3	4.0	3.8	15.0	3.3	5.1	2.8	30.6	24.4	14.9	8.7	3.1	3.2	3.6	55.9	12.7	27.5	2.35	7.4	2.3
K. indochinensis	ZMMU - A- 8153	M		48.8	11.8	4.9	4.0	3.6	15.3	3.0	5.0	2.7	31.3	24.3	14.5	7.8	2.9	3.3	3.6	53.0	12.7	26.2	2.80	6.6	2.5
K. indochinensis	ZMMU - A- 8154	M		49.8	11.9	5.5	3.9	3.5	16.3	3.4	5.0	2.7	32.0	24.3	15.1	8.4	2.7	3.1	3.8	54.5	12.9	27.5	2.62	7.4	2.5
K. indochinensis	ZMMU - A- 6316	M	topotype	50.4	11.7	5.2	4.0	3.7	14.9	3.2	4.9	2.9	28.1	22.8	14.5	8.6	3.2	3.3	3.1	54.7	12.4	27.0	2.75	6.8	2.3
K. indochinensis	ZMMU - A- 6317	M	topotype	47.3	10.8	5.1	3.8	3.3	14.1	3.0	4.9	2.9	26.7	22.0	13.9	8.0	2.7	2.9	2.8	52.2	12.2	25.7	2.89	5.9	1.8
K. indochinensis	ZMMU - A- 6318	M	topotype	51.1	11.6	5.4	3.9	3.5	16.2	3.3	5.2	2.7	31.7	25.2	15.5	8.9	3.4	3.4	3.5	55.8	13.0	27.5	2.84	7.1	2.2
K. indochinensis	ZMMU - A- 6319	M	topotype	59.6	12.7	5.8	4.1	4.0	17.7	3.4	5.8	3.1	34.3	28.6	18.1	10.4	3.9	4.4	4.2	63.9	15.0	31.6	2.79	8.5	2.8
K. indochinensis	ZMMU - A- 6320	M	topotype	51.3	12.3	5.3	4.0	3.8	15.9	3.4	5.3	2.9	30.5	24.9	15.7	9.2	3.3	3.2	3.4	55.4	13.1	27.3	2.79	7.7	2.2
K. indochinensis	ZMMU - A- 6321	M	topotype	50.0	12.3	5.3	4.0	3.7	16.1	3.4	5.4	3.2	28.4	24.0	14.7	8.8	2.9	3.4	3.4	55.7	13.4	27.5	3.22	7.0	2.1
K. indochinensis	ZMMU - A- 6322	M	topotype	50.5	12.4	6.0	4.2	4.2	16.5	3.6	5.7	3.0	30.4	24.9	15.7	9.4	3.5	3.5	3.9	57.8	13.9	28.8	2.88	7.8	2.4
K. indochinensis	ZMMU - A- 6323	M	topotype	51.7	11.4	5.4	4.0	3.7	15.0	3.6	5.9	3.3	31.3	25.3	15.8	9.2	3.6	3.9	3.2	58.0	14.1	28.0	2.85	7.4	2.4
K. indochinensis	ZMMU - A- 6324	M	topotype	50.2	12.4	5.5	4.0	3.5	15.7	2.9	5.0	3.1	29.1	23.6	14.6	8.2	3.3	3.4	3.2	55.7	13.6	26.5	2.97	7.0	2.2
K. indochinensis	ZMMU - A- 6325	M	topotype	48.2	10.7	5.1	3.9	3.7	14.3	3.3	5.1	3.0	28.8	23.1	14.4	7.8	2.7	2.9	3.4	53.9	12.4	26.3	2.65	6.7	2.1
K. indochinensis	ZMMU - A- 6315	F	topotype	55.2	13.1	6.2	4.3	4.4	17.3	3.4	5.8	3.1	32.9	27.9	18.0	9.8	3.2	3.8	4.0	62.7	14.8	31.9	3.08	8.4	2.5
K. indochinensis	ZMMU - A- 8155	SF		36.1	8.7	4.4	3.4	2.9	11.6	2.6	3.9	2.1	24.8	18.8	11.7	5.9	2.2	2.4	3.0	41.6	10.4	21.3	1.69	5.7	2.1