Unravelling cryptic species diversity in eastern Nearctic paper wasps, Polistes (Fuscopolistes), using male genitalia, morphometrics and DNA barcoding, with descriptions of two new species (Hymenoptera: Vespidae) Author Buck, Matthias Author Cobb, Tyler P. Author Stahlhut, Julie K. Author Hanner, Robert H. text Zootaxa 2012 2012-10-01 3502 1 48 journal article 44310 10.11646/zootaxa.3502.1.1 6ba8f5c9-f02f-4529-9b47-6fb31fc4a585 1175-5326 210194 6126D769-A131-49DD-B07F-0386E62FF5B9

TABLE 2. Morphometric parameters: females of Polistes hirsuticornis sp. nov., P. parametricus sp. nov. and similar species. Explanations: Avg—arithmetic mean, Range—observed minimum and maximum values (ranges marked by an asterisk include measurements that were not part of the sample used for the statistical analyses), SD—standard deviation, n —number of measured specimens, THSD—Tukey's HSD (pairwise comparisons): species with identical letters have mean values that are not statistically different (p > 0.05), F—F statistic for ANOVA within each morphometric parameter. For each ANOVA, p value was <0.001 except for CW/CL (p = 0.294; no statistically significant differences).

Morphometric parameter—Ƥ Species	Avg	Range	SD n	THSD	F
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Clypeal width / length metricus (CW/CL) bellicosus	0.98 0.99	0.91-1.04 0.95-1.04	0.029 32 0.023 25
fuscatus	0.99	0.94-1.03	0.024 24		1.25
parametricus	0.99	0.94-1.03*	0.025 25
hirsuticornis	1.01	0.96-1.06	0.025 25
Clypeus-eye contact length / Antennal hirsuticornis socket vertical diameter parametricus (CEC/AS) bellicosus	0.61 0.74 0.85	0.38-0.85 0.54-0.97 0.65-1.08	0.09 25 0.11 25 0.09 25	a b c	75.19
metricus	1.02	0.86-1.21	0.11 32	d
fuscatus	0.99	0.76-1.23*	0.11 25	d
Flagellomere 1 length / bellicosus Scape length fuscatus (FIL/SL) parametricus	0.74 0.75 0.76	0.69-0.77 0.68-0.79* 0.71-0.83	0.021 25 0.019 25 0.024 25	a b b	15.30
hirsuticornis	0.77	0.74-0.79	0.014 25	bc
metricus	0.78	0.73-0.82	0.022 32	c
Genal width / metricus Eye width bellicosus (GW/EW) fuscatus	0.73 0.76 0.78	0.48-0.85 0.67-0.90 0.67-0.92*	0.067 32 0.055 25 0.060 25	a ab bc	56.14
hirsuticornis	0.81	0.69-0.90	0.054 25	c
parametricus	0.99	0.80-1.21	0.095 25	d
Postocellar line / parametricus Ocellocular line metricus (POL/OOL) fuscatus	0.47 0.49 0.51	0.38-0.58* 0.43-0.56 0.44-0.65*	0.031 25 0.035 32 0.032 25	a ab bc	48.59
bellicosus	0.53	0.48-0.63	0.037 25	c
hirsuticornis	0.60	0.51-0.69	0.046 25	d
Scape length / width parametricus (SL/SW) hirsuticornis	3.28 3.28	3.15-3.43 2.94-3.64	0.08 25 0.15 25	a a
bellicosus	3.51	3.26-3.72	0.10 25	b	69.36
fuscatus	3.53	3.35-3.69	0.09 25	b
metricus	3.64	3.43-3.79	0.09 32	c
Upper interocular distance / metricus Head width fuscatus (UID/HW) bellicosus	0.42 0.43 0.43	0.40-0.44 0.41-0.44 0.42-0.45	0.009 32 0.008 25 0.008 25	a b b	90.18
parametricus	0.44	0.42-0.47	0.012 25	c
hirsuticornis	0.46	0.45-0.48	0.009 25	d