Unravelling cryptic species diversity in eastern Nearctic paper wasps, Polistes (Fuscopolistes), using male genitalia, morphometrics and DNA barcoding, with descriptions of two new species (Hymenoptera: Vespidae) Author Buck, Matthias Author Cobb, Tyler P. Author Stahlhut, Julie K. Author Hanner, Robert H. text Zootaxa 2012 2012-10-01 3502 1 48 journal article 44310 10.11646/zootaxa.3502.1.1 6ba8f5c9-f02f-4529-9b47-6fb31fc4a585 1175-5326 210194 6126D769-A131-49DD-B07F-0386E62FF5B9

TABLE 4. Male genitalia: morphometric parameters of aedeagus, digitus and paramere of eastern Nearctic Fuscopolistes. Explanations: Avg—arithmetic mean, Range—observed minimum and maximum values, SD—standard deviation, n —number of measured specimens. P. bellicosus WA is a form with wide aedeagus that might be a different species than the typical form. " Polistes carolina " possesses two different forms that correspond to separate clusters in the genetic analysis (see discussion of molecular study).

Aedeagus: bellicosus WA Length from base of median fuscatus expansion to apex / Width of median expansion (AL/MEW) carolina A metricus	1.61 1.68 1.80 1.82	1.54–1.67 0.09 1.56–1.82 0.09 1.69–1.88 0.10 1.69–1.94 0.12	2 10 3 5	AR, FL AR, FL, MD, TX, ON AR, SC FL, MD, MO, TX
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bellicosus	2.00	1.88–2.11 0.10	6	FL, MD, TX
rubiginosus	2.14	2.00–2.22 0.08	5	MO, TX
parametricus	2.17	2.14–2.22 0.05	3	MD
hirsuticornis	2.24	2.08–2.50 0.22	3	MD
carolina X/TX	2.24	2.11–2.40 0.11	5	FL, TX
dorsalis	2.63	2.38–3.00 0.23	5	FL, MD, TX
Aedeagus: bellicosus WA Length from base of median metricus expansion to apex / carolina A Minimum width near apex (AL/MPW) fuscatus bellicosus	5.1 5.3 5.5 5.5 7.1	4.8–5.5 0.46 5.0–6.0 0.38 5.0–6.0 0.50 4.4–6.3 0.59 6.1–8.3 0.78	2 5 3 10 6	as above
hirsuticornis	7.6	6.3–10.0 1.65	3
rubiginosus	8.3	7.1–10.0 1.18	5
parametricus	8.9	8.0–10.0 1.03	3
carolina X/TX	9.6	8.3–11.5 1.27	5
dorsalis	11.5	10.0–15.4 2.24	5
Aedeagus maximum tooth length (μm) dorsalis rubiginosus hirsuticornis carolina A	35.0 36.8 38.5 42.8	32–40 3.3 32–40 3.4 32–40 4.7 40–47 3.4	5 5 3 4
bellicosus WA	43.5	42–45 2.1	2	as above
parametricus	44.0	40–48 4.0	3
carolina X/TX	44.6	39–48 4.0	5
bellicosus	45.5	42–48 2.5	6
metricus	48.4	44–50 3.1	5
fuscatus	71.0	64–88 10.5	10
Digitus: fuscatus Apical process length / Total length metricus bellicosus	0.17 0.18 0.21	0.10–0.21 0.032 0.15–0.20 0.021 0.18–0.24 0.026	10 5 6
carolina A	0.22	0.21–0.24 0.013	4
bellicosus WA	0.23	0.21–0.24 0.021	2	as above
parametricus	0.23	0.21–0.27 0.038	3
carolina X/TX	0.23	0.19–0.28 0.032	5
hirsuticornis	0.24	0.21–0.26 0.021	3
rubiginosus dorsalis	0.26 0.26	0.23–0.29 0.024 0.24–0.28 0.018	5 5