Two new species and a new record of Helvella (Helvellaceae, Pezizales) from South Korea Author Park, Sang-Young 0009-0008-5453-8902 Department of Forest Science, Chungbuk National University, Cheongju, 28644, Republic of Korea fungux@naver.com Author Kim, Sohee 0009-0006-5083-4451 Department of Forest Science, Chungbuk National University, Cheongju, 28644, Republic of Korea kha1480@naver.com Author Park, Eunsu 0000-0003-4605-3661 Ecological Technology Research Team, Division of Ecological Applications Research, National Institute of Ecology, Seocheon, 33657, Republic of Korea eunsu156@naver.com Author Kwag, Young-Nam 0000-0001-7333-2303 Forest Biodiversity Division, Korea National Arboretum, Pocheon 11186, Republic of Korea kyn0102@korea.kr Author Eo, Ju-Kyeong 0000-0001-8376-5575 National Ecological Survey Team, Division of Ecological Survey, National Institute of Ecology, Seocheon, 33657, Republic of Korea abiesendo@gmail.com Author Lee, Hwayong 0000-0003-4526-2082 Department of Forest Science, Chungbuk National University, Cheongju, 28644, Republic of Korea leehy@chungbuk.ac.kr text Phytotaxa 2025 2025-12-04 732 2 147 162 https://doi.org/10.11646/phytotaxa.732.2.3 journal article 314347 10.11646/phytotaxa.732.2.3 e0639f70-03e5-40ba-b854-deb313d99f41 1179-3163 18423175

TABLE 1. Korean Helvella species and sequences analyzed in this study. Species names in bold indicate sequences obtained from the ten specimens that were examined for both morphological and molecular phylogenetic analyses. A hyphen (–) represents data not available.

Species name	Type	Sample ID	Voucher No.	Accession no.			Country
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				ITS	LSU	hsp90
Dissingia oblongispora			HSA 137	OR355181	OR355355	OR366160	China
Dissingia oblongispora			HSA 169	OR355182	OR355356	OR366161	China
Helvella acetabuloides	holotype		CFSZ2044	OR355015	OR355185	MK652219	China
Helvella acetabuloides			BJTC FM1264	OR355166	OR355337	OR366140	China
H. acetabulum		H134	O-64925	-	KY772985	KY784267	Norway
H. acetabulum		H225	O-253212	-	KY773055	KY784344	Norway
H. acetabulum	epitype	H410	C-F Fungi Exs. Suec. 1354	-	-	KY784506	Sweden
H. acetabulum		KH089	KA25-0481	PV830655	PV830661	PV853822	Korea
H. alborava	holotype		BJTC C316-A	OR355106	OR355276	OR366098	China
H. alborava			BJTC C316-B	OR355173	OR355347	OR366148	China
H. arctoalpina			N127	OQ065745	PP396828	-	Turkey
H. arctoalpina			VANF127-2	PQ512823	PQ507477	-	Turkey
H. arctoalpina			VANF127-3	PQ512824	PQ507478	-	Turkey
H. atroides			MFLU23-0101	-	OQ866058	OQ863540	Thailand
H. atroides	holotype		MFLU23-0102	-	OQ866059	OQ863541	Thailand
H. atroides			MFLU23-0103	-	OQ866060	OQ863542	Thailand
H. elastica		H066	O-253311	-	KY772950	KY784230	Sweden
H. fistulosa			MFLU23-0096	-	OQ866053	OQ863535	Thailand
H. fistulosa			MFLU23-0097	-	OQ866054	OQ863536	Thailand
H. fistulosa			MFLU23-0098	-	OQ866055	OQ863537	Thailand
H. griseomacropus sp. nov.		H41	KA16-0551	PP545146	PP544953	-	Korea
H. griseomacropus sp. nov.		H59	KA20-0060	PP545162	PP544970	-	Korea
H. griseomacropus sp. nov.		H72	KA21-1232	PP545173	PP544981	-	Korea
H. griseomacropus sp. nov.		KH027	KA24-1348	PP545244	PP545067	PP782836	Korea
H. griseomacropus sp. nov.		KH028	KA24-1349	PP545245	PP545068	PP782837	Korea
H. griseomacropus sp. nov.	holotype	KH099	KA25-0485	-	PV830665	PV853826	Korea
H. griseomacropus sp. nov.		N39	NIBRFG0000511082	PP545198	PP545015	-	Korea
H. griseomacropus sp. nov.		N40	NIBRFG0000511110	PP545199	PP545016	-	Korea
H. griseomacropus sp. nov.		N48	NIBRFG0000513941	PP545205	PP545022	-	Korea
H. koreana sp. nov.	holotype	KH087	KA25-0479	PV830653	PV830659	PV853820	Korea
H. koreana sp. nov.		KH088	KA25-0480	PV830654	PV830660	PV853821	Korea
H. macropus		H238	O-291425	-	KY773067	KY784356	Norway
				ITS	LSU	hsp90
H. macropus		H239	O-291391	-	KY773068	-	Norway
H. macropus	epitype	H412	C-F Fungi Exs. Suec. 3266	-	-	KY784507	Sweden
H. neofistulosa	holotype		BJTC ZH1213	OR355059	OR355229	OR366056	China
H. neofistulosa			BJTC ZH1251	OR355060	OR355230	OR366057	China
H. nigrorava	holotype		BJTC ZH67	OR355105	OR355275	OR366097	China
H. orentitomentosa	holotype		MFLU23-0099	-	OQ866056	OQ863538	Thailand
H. orentitomentosa			MFLU23-0100	-	OQ866057	OQ863539	Thailand
H. pakistanica	holotype		M.jabba01	PQ410250	-	-	Pakistan
H. pakistanica			M.jabba02	PQ410251	-	-	Pakistan
H. pseudoelastica		KH056	KA24-1347	-	PP545094	PP782860	Korea
H. pseudoelastica		KH090	KA25-0482	PV830656	PV830662	PV853823	Korea
H. pseudoelastica	holotype		BJTC C351-A	OR355170	OR355341	OR366143	China
H. pseudoelastica			BJTC C351-B	OR355174	OR355348	OR366149	China
H. rivularis		H1978	O-255764	MN656175	MN655850	MN692371	Norway
H. rivularis	holotype	H276	C-F-59447	-	-	KY784391	Norway
H. rivularis			VANF714	PP359562	PP396830	-	Turkey
H. rivularis			VANF714-2	PQ512821	PQ507475	-	Turkey
H. rivularis			VANF714-3	PQ512822	PQ507476	-	Turkey
H. rugosa			MFLU23-0093	-	OQ866050	OQ863532	Thailand
H. rugosa			MFLU23-0094	-	OQ866051	OQ863533	Thailand
H. rugosa			MFLU23-0095	-	OQ866052	OQ863534	Thailand
H. solitaria		H090	O-253371	-	KY772959	KY784241	France
H. solitaria		H215	O-253379	-	KY773047	KY784336	Norway
H. solitaria			BJTC FM417	OR355131	OR355302	OR366120	China
H. sp. K18		KH092	KA25-0483	PV830657	PV830663	PV853824	Korea
H. sp. K7		KH096	KA25-0484	PV830658	PV830664	PV853825	Korea
H. sp. K7		S07	SFC20150813-12	PP545213	PP545030	-	Korea
H. sublactea			VANF141	PP359561	PP396829	-	Turkey
H. sublactea			VANF141-2	PQ512819	PQ507473	-	Turkey
H. sublactea			VANF141-3	PQ512820	PQ507474	-	Turkey
H. sublicia		H148	O-70080	-	KY772997	KY784281	Norway
H. sublicia			KH.11.84(S)	-	MK100273	MK179403	Sweden
H. submacropus	holotype		HKAS 70217	KX506977	KX506932	OR220563	China
H. submacropus			HKAS 78944	-	KX506934	OR220561	China
H. submacropus			HKAS 78930	KX506978	KX506933	OR220562	China
H. submacropus			HKAS 57739	KX506980	KX506935	OR220560	China
H. submacropus			HKAS 90593	KX506982	KX506937	OR220559	China
H. taiyuanensis			HMAS 277500	MK592152	MK592105	MK652216	China
H. taiyuanensis			119250	MK592151	MK592104	MK652215	China
H. taiyuanensis			BJTC FM192	OR355167	OR355338	OR366141	China
H. taiyuanensis			BJTC FM861	OR355168	OR355339	OR366142	China
H. ulvinenii			HKAS 90321	-	KU167483	OR366155	China