Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae) Author Vinarski, Maxim V. Laboratory of Macroecology and Biogeography of Invertebrates, Saint-Petersburg State University, 7 / 9 Universitetskaya Emb., 199034, Saint-Petersburg, Russia & Omsk State University, 28 Adrianova Str., 644077, Omsk, Russia https://orcid.org/0000-0002-7644-4164 radix.vinarski@gmail.com Author Aksenova, Olga V. Laboratory of Macroecology and Biogeography of Invertebrates, Saint-Petersburg State University, 7 / 9 Universitetskaya Emb., 199034, Saint-Petersburg, Russia & N. Laverov Federal Center for Integrated Arctic Research, Ural Branch of the Russian Academy of Sciences, 23 Severnaya Dvina Emb., 163000, Arkhangelsk, Russia https://orcid.org/0000-0002-0817-7105 Author Bolotov, Ivan N. N. Laverov Federal Center for Integrated Arctic Research, Ural Branch of the Russian Academy of Sciences, 23 Severnaya Dvina Emb., 163000, Arkhangelsk, Russia & Northern (Arctic) Federal University, 17 Severnaya Dvina Emb., 163002, Arkhangelsk, Russia https://orcid.org/0000-0002-3878-4192 text Zoosystematics and Evolution 2020 96 2 577 608 http://dx.doi.org/10.3897/zse.96.52860 journal article http://dx.doi.org/10.3897/zse.96.52860 1860-0743-2-577 D4882E78D81B5FA394A6A156886835E4

Table 1. Morphological characterisation of shells and copulative apparatuses of several species in the genus Radix.

Character / index	Species / locality
	R. alticola		R. euphratica		R. brevicauda	R. plicatula		R. natalensis	R. rufescens
	Tajikistan, Zor-Kul Lake (paratypes)	Tajikistan, a warm brook near Dzhaushangoz village*	Tajikistan, Dushanbe, a fountain near President's palace*	Tajikistan, a roadside ditch near Kurban-Shakhid village*	Tajikistan, a warm spring near Shaimak village	China, Bejing, a pond near the former Emperor's palace*	China, Uyghuria, Bagrashkol' Lake*	Uganda, the Kyamwiga crater lake*, **	India, "Khipru, Sind"	Myanmar, Yetho River near the dam*
*Sample studied genetically. ** The short-spired morph of R. natalensis.
Repository	ZISP	LMBI	LMBI	LMBI	ZISP	LMBI	LMBI	LMBI	NHMUK	LMBI
Number of specimens measured (dissected)	45 (0)	44 (16)	4(3)	21(15)	21(0)	29(11)	16(9)	26(20)	40(0)	12(6)
Whorls number	3.00-3.75	3.00-3.75	3.87-4.25	3.25-4.12	3.25-4.00	3.75-4.00	3.50-4.12	3.25-3.75	3.75-4.75	3.87-4.50
	3.40 +/- 0.18	3.30 +/- 0.17	4.06 +/- 0.16	3.71 +/- 0.21	3.48 +/- 0.19	4.08 +/- 0.21	3.76 +/- 0.23	3.51 +/- 0.15	4.17 +/- 0.21	4.22 +/- 0.22
Shell height, mm (SH)	9.0-13.9	9.4-15.3	12.6-15.1	12.5-20.0	9.8-15.3	13.6-20.7	12.9-22.6	13.6-18.8	37.9-43.4	17.5-28.9
	10.9 +/- 1.0	11.9 +/- 1.4	13.9 +/- 1.1	14.8 +/- 2.1	12.1 +/- 1.5	16.5 +/- 1.5	15.8 +/- 3.1	16.2 +/- 1.3	39.9 +/- 1.6	23.2 +/- 3.8
Shell width, mm (SW)	6.4-10.5	7.8-13.2	8.4-9.3	7.6-13.4	8.4-13.0	7.9-14.2	8.2-14.6	8.0-13.0	17.8-20.8	9.1-13.6
	8.1 +/- 0.8	9.5 +/- 1.2	8.9 +/- 0.4	9.7 +/- 1.5	10.3 +/- 1.4	10.4 +/- 1.6	10.1 +/- 2.1	10.5 +/- 1.0	18.8 +/- 1.0	11.0 +/- 1.4
Spire height, mm (SpH)	1.4-2.8	2.0-4.0	3.8-4.5	2.7-5.2	2.2-4.2	3.4-6.8	2.7-7.0	2.8-4.7	10.6-14.8	4.5-10.8
	2.1 +/- 0.3	2.6 +/- 0.4	4.1 +/- 0.3	3.7 +/- 0.8	2.8 +/- 0.5	5.1 +/- 0.9	4.4 +/- 1.2	3.7 +/- 0.5	12.7 +/- 1.1	7.7 +/- 1.8
Body whorl height, mm (BWH)	8.4-12.8	8.7-14.2	10.1-12.2	11.0-17.6	8.9-13.8	11.7-18.0	11.5-20.3	12.3-17.5	32.8-37.4	15.6-25.2
	9.9 +/- 0.9	10.9 +/- 1.3	11.7 +/- 1.1	13.1 +/- 1.9	10.9 +/- 1.4	14.4 +/- 1.5	14.1 +/- 2.8	14.8 +/- 1.2	34.7 +/- 1.2	20.1 +/- 3.3
Aperture height, mm (AH)	7.6-11.4	7.7-12.9	9.1-12.2	9.4-15.1	7.7-11.7	8.8-16.0	9.4-16.1	10.2-14.6	27.0-31.2	12.7-19.1
	9.1 +/- 0.9	9.6 +/- 1.2	10.1 +/- 0.9	11.4 +/- 1.6	9.6 +/- 1.2	11.8 +/- 1.7	11.6 +/- 2.2	12.6 +/- 1.0	28.9 +/- 1.2	15.5 +/- 2.2
Aperture width, mm (AW)	5.0-8.5	5.2-10.4	5.8-7.3	5.8-11.8	5.5-8.8	6.3-12.4	6.1-10.5	6.0-9.3	12.3-16.3	6.5-11.3
	6.3 +/- 0.7	6.4 +/- 1.1	6.5 +/- 0.6	7.5 +/- 1.5	7.0 +/- 1.0	8.2 +/- 1.4	7.5 +/- 1.6	7.9 +/- 0.7	14.4 +/- 0.9	8.6 +/- 1.4
Praeputium length, mm (PP)	-	3.2-5.6	2.8-3.6	3.2-4.9	-	3.4-6.5	3.0-5.4	3.1-4.9	-	2.8-4.2
		4.1 +/- 0.6	3.2 +/- 0.9	3.9 +/- 1.1		4.3 +/- 0.9	4.2 +/- 0.9	4.3 +/- 0.5		3.4 +/- 1.0
Penis sheath length, mm (PS)	-	3.2-5.1	2.9-3.5	2.2-3.9	-	3.6-9.2	3.6-7.8	3.0-5.5	-	2.2-3.6
		4.2 +/- 0.6	3.2 +/- 0.6	3.1 +/- 1.1		5.8 +/- 1.2	5.5 +/- 1.3	4.5 +/- 0.6		2.9 +/- 1.0
SW/SH	0.65-0.81	0.72-0.87	0.62-0.67	0.61-0.73	0.79-0.94	0.53-0.76	0.55-0.72	0.57-0.71	0.44-0.50	0.41-0.52
	0.74 +/- 0.03	0.80 +/- 0.04	0.64 +/- 0.03	0.66 +/- 0.03	0.85 +/- 0.04	0.63 +/- 0.05	0.64 +/- 0.04	0.65 +/- 0.04	0.47 +/- 0.02	0.48 +/- 0.03
SpH/SH	0.13-0.24	0.18-0.27	0.28-0.32	0.20-0.30	0.18-0.28	0.20-0.42	0.20-0.33	0.19-0.27	0.27-0.38	0.26-0.37
	0.19 +/- 0.02	0.22 +/- 0.02	0.30 +/- 0.02	0.25 +/- 0.03	0.23 +/- 0.02	0.31 +/- 0.05	0.28 +/- 0.04	0.23 +/- 0.02	0.32 +/- 0.02	0.33 +/- 0.04
BWH/SH	0.87-0.93	0.88-0.98	0.80-0.86	0.83-0.94	0.87-0.92	0.83-0.95	0.86-0.92	0.86-0.93	0.84-0.89	0.80-0.90
	0.90 +/- 0.01	0.91 +/- 0.02	0.84 +/- 0.03	0.89 +/- 0.02	0.90 +/- 0.01	0.87 +/- 0.02	0.89 +/- 0.02	0.91 +/- 0.02	0.87 +/- 0.01	0.86 +/- 0.02
AH/SH	0.65-0.81	0.75-0.85	0.70-0.74	0.70-0.86	0.72-0.86	0.59-0.86	0.69-0.81	0.74-0.82	0.69-0.77	0.63-0.75
	0.74 +/- 0.03	0.81 +/- 0.02	0.73 +/- 0.02	0.77 +/- 0.04	0.80 +/- 0.03	0.71 +/- 0.06	0.74 +/- 0.04	0.78 +/- 0.02	0.73 +/- 0.02	0.67 +/- 0.04
AW/AH	0.59-0.83	0.66-0.91	0.63-0.67	0.57-0.78	0.67-0.81	0.63-0.78	0.56-0.73	0.57-0.67	0.45-0.55	0.50-0.61
	0.69 +/- 0.04	0.72 +/- 0.04	0.65 +/- 0.02	0.65 +/- 0.05	0.73 +/- 0.03	0.70 +/- 0.04	0.64 +/- 0.05	0.62 +/- 0.02	0.50 +/- 0.03	0.55 +/- 0.04
ICA	-	0.81-1.18	0.93-1.16	1.10-1.69	-	0.54-0.96	0.64-0.90	0.84-1.13	-	1.09-1.27
		0.97 +/- 0.11	1.01 +/- 0.12	1.25 +/- 0.18		0.76 +/- 0.14	0.79 +/- 0.08	0.96 +/- 0.07		1.19 +/- 0.06