Euroscaptor darwini sp. nov., a new species of mole (Mammalia, Eulipotyphla, Talpidae) from the north-central mountains in Vietnam Author Nguyen, Son Truong 0000-0003-3214-4407 Institute of Biology, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet Road, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet Road, Hanoi, Vietnam Author Bui, Hai Tuan 0000-0003-4065-7229 Faculty of Biology, Hanoi University of Science, Vietnam National University (VNU), 334 Nguyen Trai Road, Hanoi, Vietnam Author Dau, Vinh Quang 0000-0002-8976-1644 Hong Duc University, 565 Quang Trung Road, Thanh Hoa, Vietnam Author Le, Phuong Dinh Pu Luong Nature Reserve, Thanh Hoa, Vietnam Author Vu, Yen Huong https://orcid.org/0009-0001-4643-167X Institute of Biology, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet Road, Hanoi, Vietnam & Faculty of Biology, Hanoi University of Science, Vietnam National University (VNU), 334 Nguyen Trai Road, Hanoi, Vietnam text ZooKeys 2025 2025-10-10 1255 239 274 journal article 312575 10.3897/zookeys.1255.161942 1abe6cfb-401a-4d86-94e7-6b3023e65ed2 3C3E0CF2-85BF-4F5C-B4D3-9140F0BE2BC9

Table 3. Comparative statistics of external measurements (in millimeters) between E. darwini sp. nov. and other fossorial mole species. (*) indicates shared measurements between E. parvidens and E. ngoclinhensis; (#) indicates shared measurements between E. kuznetsovi and E. orlovi.

Species	Reference (n)	HB	T	HF	FFW	FFL	Tail ratio = T / HB (%)
E. darwini sp. nov.	This study (holotype ♂)	115.71	2.11	14.51	13.88	13.29	1.82
	This study (4 paratypes ♀)	122.21–127.93 125.52 ± 2.86	2.67–3.74 3.33 ± 0.47	14.86–15.38 15.03 ± 0.24	14.02–14.51 14.21 ± 0.22	13.64–14.11 13.86 ± 0.28	2.18–2.92 2.64 ± 0.32
	This study (both sexes)	115.71–127.93 123.56 ± 5.04	2.11–3.74 3.08 ± 0.68	14.51–15.38 14.93 ± 0.31	13.88–14.51 14.14 ± 0.24	13.29–14.11 13.79 ± 0.41	1.82–2.92 2.48 ± 0.46
E. subanura	Kawada et al. (2012) (8)	113.0–130.0 122.88 ± 5.51	4.0–5.0 4.38 ± 0.44	13.5–15.5 14 ± 0.38	15.0–16.5 14.38 ± 0.38	14.0–15.5 14.75 ± 0.46	3.6
	Kawada (2016) (28)	113.0–131.5 125.48 ± 3.95	2.5–5.0 3.80 ± 0.79	13.0–14.5 13.70 ± 0.49	13.0–15.5 14.21 ± 0.55	13.0–15.5 14.27 ± 0.62	1.96–4.24 3.04 ± 0.65
	Bui (2022) (42)	107.0–130.5 121.89 ± 5.61	2.50–5.00 3.76 ± 0.73	12.0–16.5 14.02 ± 0.87	13.0–15.5 14.29 ± 0.55	13.80–18.00 14.80 ± 0.84	2.33–3.85 3.08 ± 0.56
E. parvidens	Kawada et al. (2012) (10) *	135.0–148.5 143.75 ± 3.75	5.5–9.0 6.95 ± 1.09	13.5–15.5 14.6 ± 0.66	15.0–16.5 15.4 ± 0.61	14.5–16.5 15.3 ± 0.63	4.8
	Kawada (2016) (18) *	135.0–150.0 144.89 ± 3.77	5.5–10.5 7.42 ± 1.30	13.5–16.0 14.92 ± 0.69	14.0–17.5 15.47 ± 0.88	14.5–18.0 15.75 ± 0.93	4.07–7.07 5.11 ± 0.83
	Bui (2022) (42)	130–165.5 150.9 ± 10.71	5.5–10.0 7.9 ± 1.48	12.0–17.74 16.01 ± 1.18	14.15–16.96 15.65 ± 0.54	14.7–17.6 16.03 ± 0.80	4.23–6.06 5.23 ± 0.0. 75
E. ngoclinhensis	Bui (2022) (30)	120.0–150.0 137.87 ± 9.01	5.50–10.0 7.50 ± 1.18	12.0–16.0 14.15 ± 0.93	12.27–17.0 14.66 ± 1.34	13.20–18.00 15.02 ± 1.09	4.58–6.67 5.44 ± 0.86
E. kuznetsovi	Kawada et al. (2012) (19) #	133.0–144.0 138.45 ± 3.39	14.0–17.0 15.37 ± 1.01	14.5–16.5 15.55 ± 0.50	14.5–17.0 15.95 ± 0.71	15.0–17.5 16.26 ± 0.77	11.1
	Kawada (2016) (27) #	122.0–144.0 136.85 ± 4.97	10.0–17.0 14.69 ± 2.00	14.5–16.5 15.46 ± 0.46	14.5–17.0 15.91 ± 0.72	15.0–17.5 16.24 ± 0.74	8.19–11.81 10.73 ± 0.93
	Zemlemerova et al. (2016)	132–136					11.4–12.5
	Kryštufek and Motokawa (2018)	117–136	14–17	14.5–16.5			11.6–14.5
	Bui (2022) (24)	115.4–144 137.37 ± 5.74	14–17.5 15.65 ± 1.09	14.5–18.4 15.73 ± 0.77	14.5–17 15.82 ± 0.69	15–17.9 16.36 ± 0.80	11.33–12.57 12.05 ± 0.4
E. orlovi	Zemlemerova et al. (2016)	115–129					12.6–13.7
	Kryštufek and Motokawa (2018)	106–129	15–17.5	15.5			12.5–15.1
	Bui (2022) (6)	115–137.62 127 ± 7.40	11.76–19.28 16 ± 2.29	13.43–15.5 14.87 ± 0.92	14–16.5 15.15 ± 0.73	15–16.5 15.83 ± 0.76	10.63–13.57 12.45 ± 0.67
E. longirostris	Smith and Xie (2008)	90–145	11.0–25	14–23
E. klossi	Kawada et al. (2012) (2)	130.0–135.5	9.5–10.5	15	16	16.0–16.5	7.5
	Kawada (2016) (4)	115.0–135.5 127.63 ± 8.81	9.5–13 11.25 ± 1.55	15.0–16.0 15.38 ± 0.48	16.0–17.0 16.38 ± 0.48	15.5–16.5 16.13 ± 0.48	7.01–11.30 8.91 ± 1.84
	Smith and Xie (2008)	115–135.5	9.5–13	15–16			7–11.3
E. malayana	Kawada et al. (2008)	124–134.5	4.5–8.5	14–16.5	15–17	15–19	3.35–6.59
	Kawada (2016) (10)	128.5–134.5 131.15 ± 1.94	4.5–8.5 5.70 ± 1.09	14.5–16.5 15.50 ± 0.62	15.5–17.0 16.40 ± 0.57	15.0–16.5 15.80 ± 0.42	3.34–6.58 4.35 ± 0.86
	Smith and Xie (2008)	128.5–134.5	4.5–8.5	14.5–16.5			3.3–6.6
E. grandis	Miller (1940)	150	9.6	18	15
E. micrurus	Smith and Xie (2008)	128–135	5–9	15–16